Drymocallis arizonica
Drymocallis lactea
Arizona drymocallis or wood beauty
Nevada cinquefoil, Sierran woodbeauty
short.
short.
± tufted, (1.5–)2.5–6 dm;
base 1.5–2.5 mm diam., sparsely to densely septate-glandular.
tufted, (0.3–)1.5–6(–6.5) dm;
base 1–2.5 mm diam., not or sparsely, sometimes moderately, septate-glandular.
sparsely to moderately hairy;
basal 5–22 cm, leaflet pairs (2–)3–4(–5);
terminal leaflet broadly obovate-elliptic, 1–5 × 1–2.5(–4) cm, teeth double, 6–18 per side, apex rounded to acute;
cauline 1–4, well developed, leaflet pairs 2–3.
usually sparsely to moderately hairy, sometimes glabrate;
basal (2–)5–20(–30) cm, leaflet pairs (2–)3–4(–5);
terminal leaflet broadly obovate to cuneate, 1–4 × 0.7–3.5 cm, teeth usually single, 4–10(–14) per side, apex usually rounded to obtuse, rarely acute;
cauline 1–2, reduced, leaflet pairs 2–3.
5–30-flowered, not notably leafy, compact, 1/10–1/4(–1/2) of stem, narrow, branch angles 5–25°.
(2–)5–30(–50)-flowered, not leafy, open, (1/6–)1/5–1/2(–4/5) of stem, narrow to wide, branch angles 10–40(–50)°.
1–10 (proximal to 20) mm, sparsely to ± densely short-hairy, predominantly septate-glandular.
2–10 (proximal to 30) mm, predominantly short-hairy, sometimes velutinous, not or sparsely, sometimes moderately, septate-glandular.
opening widely;
epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–4(–7) × 1–2 mm;
sepals spreading, 4–6(–11) mm, apex ± acute;
petals not or scarcely overlapping, spreading, cream-white, ± obovate, 3–6 × 2.5–5 mm, shorter than or equal to sepals;
filaments 1–3 mm, anthers 0.7–1 mm;
styles thickened, 1 mm.
opening widely;
epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm;
sepals spreading, 3–8(–9) mm, apex usually acute, sometimes obtuse and apiculate;
petals ± overlapping, spreading, cream-white to bright yellow, broadly obovate, (3–)4–8(–10) × 3–8 mm, usually longer than, rarely equal to, sepals;
filaments 1–3(–3.5) mm, anthers 0.6–1 mm;
styles thickened, 1 mm.
light brown, 1 mm.
light brown, 1 mm.
= 14.
Drymocallis arizonica
Drymocallis lactea
Drymocallis arizonica encompasses populations in Arizona north of the Mogollon Rim and in Utah as far north as Garfield and Sevier counties, including the Henry Mountains. It is most distinctive in northern Arizona, where plants commonly have basal leaves with four pairs of lateral leaflets and compact inflorescences. The species intergrades with D. convallaria but is in general shorter and more likely multistemmed. It also shares similarities with D. pseudorupestris but has more developed cauline leaves and a more compact inflorescence. The intergrade zone between all three species and D. deseretica in southwestern Utah is particularly problematic and perhaps indicative of yet additional taxa deserving recognition.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Varieties 2 (2 in the flora).
Drymocallis lactea has had a well-established identity as Potentilla glandulosa var. (or subsp.) nevadensis; however, the epithet lactea has priority at species rank and avoids possible confusion with P. nevadensis Boissier. D. D. Keck (in J. Clausen et al. 1940) based his concept of P. glandulosa subsp. nevadensis on white-petaled plants with narrow inflorescences, the common form in the central Sierra Nevada and south. His map and annotations, however, include populations with yellow petals and widely branched inflorescences from northern California and adjacent Oregon. The two extremes merge in the northern Sierra Nevada, and populations in the North Coast Ranges of California also are problematic; the extremes are accordingly treated here as varieties of a single species. The inclusion by Keck of Washington in the species range might have been based on specimens here assigned to P. pseudorupestris.
Drymocallis lactea can be distinguished from sympatric species by its relatively large, overlapping petals, non-leafy inflorescences, and singly toothed leaflets. Pedicels and distal stems are usually densely short-hairy with relatively few or no glandular hairs (in contrast to D. pseudorupestris), and septate glands are usually absent from stem bases (in contrast to D. hansenii). The distinction between D. lactea and D. pseudorupestris breaks down at the margin of the species range in Nevada and Oregon.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Inflorescences narrow, branch angles 10–20°; petals cream-white to pale yellowish; pedicels and stems not or sparsely septate-glandular. | var. lactea |
1. Inflorescences wide, branch angles mostly 20–40°; petals usually pale to bright yellow; pedicels and stems sometimes moderately septate-glandular. | var. austiniae |
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