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Arizona drymocallis or wood beauty

Douglas' woodbeauty, Greene's drymocallis, sticky cinquefoil

Caudex branches

short.

short.

Stems

± tufted, (1.5–)2.5–6 dm;

base 1.5–2.5 mm diam., sparsely to densely septate-glandular.

tufted, (1–)1.5–7.5 dm;

base 1–5 mm diam., moderately to densely septate-glandular.

Leaves

sparsely to moderately hairy;

basal 5–22 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate-elliptic, 1–5 × 1–2.5(–4) cm, teeth double, 6–18 per side, apex rounded to acute;

cauline 1–4, well developed, leaflet pairs 2–3.

sparsely to moderately hairy;

basal (3–)6–30 cm, leaflet pairs (1–)2–4;

terminal leaflet broadly obovate to rhombic-elliptic, (1–)1.5–7(–8) × 1–5 cm, teeth usually double (irregularly single in var. viscida), (4–)6–18(–20) per side, apex rounded to obtuse;

cauline 0–3, developed or reduced, leaflet pairs 1–4.

Inflorescences

5–30-flowered, not notably leafy, compact, 1/10–1/4(–1/2) of stem, narrow, branch angles 5–25°.

(2–)5–40-flowered, leafy or not, open or of congested clusters terminating branches, (1/10–)1/5–3/4 of stem, usually wide (narrow in var. viscida), branch angles (15–)20–55°.

Pedicels

1–10 (proximal to 20) mm, sparsely to ± densely short-hairy, predominantly septate-glandular.

1–10 (proximal to 30) mm, not or sparsely short-hairy, predominantly septate-glandular.

Flowers

opening widely;

epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–4(–7) × 1–2 mm;

sepals spreading, 4–6(–11) mm, apex ± acute;

petals not or scarcely overlapping, spreading, cream-white, ± obovate, 3–6 × 2.5–5 mm, shorter than or equal to sepals;

filaments 1–3 mm, anthers 0.7–1 mm;

styles thickened, 1 mm.

opening widely;

epicalyx bractlets linear-lanceolate to elliptic, 1.5–7(–8) × 0.5–2.5(–3) mm;

sepals spreading or reflexed, (3–)4–11 mm, apex usually broadly obtuse with mucronate tip, sometimes acute;

petals not overlapping, spreading or reflexed, cream-white to yellow, narrowly to broadly obovate or ovate to nearly round, 2–6.5 × 1.5–5.5 mm, shorter than or equal to sepals;

filaments 0.5–2 mm, anthers 0.5–1 mm;

styles thickened, 1–1.5 mm.

Achenes

light brown, 1 mm.

reddish brown, 0.9–1.5 mm.

2n

= 14.

Drymocallis arizonica

Drymocallis glandulosa

Phenology Flowering May–Jul.
Habitat Streamsides, rocky sites, open forest floors, pine and aspen forests
Elevation 1800–3200 m (5900–10500 ft)
Distribution
from FNA
AZ; UT
[BONAP county map]
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; BC; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Drymocallis arizonica encompasses populations in Arizona north of the Mogollon Rim and in Utah as far north as Garfield and Sevier counties, including the Henry Mountains. It is most distinctive in northern Arizona, where plants commonly have basal leaves with four pairs of lateral leaflets and compact inflorescences. The species intergrades with D. convallaria but is in general shorter and more likely multistemmed. It also shares similarities with D. pseudorupestris but has more developed cauline leaves and a more compact inflorescence. The intergrade zone between all three species and D. deseretica in southwestern Utah is particularly problematic and perhaps indicative of yet additional taxa deserving recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

Even with the removal of most of the subspecies of Potentilla glandulosa treated by Keck, Drymocallis glandulosa encompasses an inordinate range of variation. This is because of the apparently clinal variation between the coastal extreme (var. wrangelliana), with leafy inflorescences and broad cream-white petals, the montane var. reflexa, with less leafy inflorescences and narrow yellow petals, and the transitional interior var. glandulosa. Variety reflexa, in turn, intergrades in the mountains of southern California with var. viscida, which is anomalous in D. glandulosa with its narrow inflorescences and irregularly single-toothed leaflets. Variation outside of California is unclear, with no evident pattern to differences in petal size and color; these have largely been assigned to var. glandulosa.

At its most distinctive, Drymocallis glandulosa is a robust, densely glandular plant with widely branched leafy inflorescences (except var. viscida), relatively short, stout pedicels, broad sepals with rounded, mucronate tips (except var. viscida and sometimes var. reflexa) that are often significantly enlarged in fruit, and non-overlapping petals that are shorter than to scarcely equal to the sepals. The relatively large achenes are blunter and redder than those of most other species of Drymocallis, as discussed by J. Clausen et al. (1940).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals obovate-elliptic to broadly ovate to nearly round, (2–)3.5–5.5 mm wide, cream-white to ± yellow; sepals spreading, apices obtuse to rounded; inflorescences ± to very leafy; pedicels 1–5 (proximal to 30) mm
→ 2
1. Petals narrowly ovate to narrowly obovate or obovate-elliptic, 1.5–3 mm wide, yellow; sepals reflexed, apices acute to obtuse; inflorescences ± leafy or not; pedicels 2–10 (proximal to 20) mm
→ 3
2. Petals (3–)3.5–5 mm, usually ± yellow, sometimes cream-white; inland.
var. glandulosa
2. Petals (4–)4.5–6.5 mm, usually cream-white, sometimes pale yellow; ± coastal.
var. wrangelliana
3. Inflorescences usually wide, branch angles (15–)25–55°; leaflet teeth usually double, rarely single.
var. reflexa
3. Inflorescences narrow, branch angles 15–30(–40)°; leaflet teeth single or double.
var. viscida
Source FNA vol. 9, p. 284. FNA vol. 9, p. 292.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa
D. arguta, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Subordinate taxa
D. glandulosa var. glandulosa, D. glandulosa var. reflexa, D. glandulosa var. viscida, D. glandulosa var. wrangelliana
Synonyms D. glandulosa subsp. arizonica, Potentilla glandulosa subsp. arizonica, P. macdougalii Potentilla glandulosa
Name authority Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 373. (1908) (Lindley) Rydberg: Monogr. N. Amer. Potentilleae, 198. (1898)
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