Drymocallis arguta
Drymocallis lactea
cordilleran drymocallis, sticky cinquefoil, tall cinquefoil, tall drymocallis or wood beauty, tall wood-beauty
Nevada cinquefoil, Sierran woodbeauty
short.
short.
usually solitary, sometimes tufted, (1–)3–10 dm;
base 3–6 mm diam., ± densely, sometimes sparsely, septate-glandular.
tufted, (0.3–)1.5–6(–6.5) dm;
base 1–2.5 mm diam., not or sparsely, sometimes moderately, septate-glandular.
usually moderately to densely hairy;
basal (6–)12–40 cm, leaflet pairs (3–)4–5;
terminal leaflet broadly elliptic-obovate, (2–)4–10 × (1–)2–4.5 cm, teeth mostly double, 15–30+ per side, apex obtuse to acute;
cauline 2–3, relatively well developed, leaflet pairs 2–4.
usually sparsely to moderately hairy, sometimes glabrate;
basal (2–)5–20(–30) cm, leaflet pairs (2–)3–4(–5);
terminal leaflet broadly obovate to cuneate, 1–4 × 0.7–3.5 cm, teeth usually single, 4–10(–14) per side, apex usually rounded to obtuse, rarely acute;
cauline 1–2, reduced, leaflet pairs 2–3.
10–40-flowered, not leafy, congested, sometimes more elongate in fruit, 1/10–1/5(–1/3) of stem, narrow, branch angles 5–20°.
(2–)5–30(–50)-flowered, not leafy, open, (1/6–)1/5–1/2(–4/5) of stem, narrow to wide, branch angles 10–40(–50)°.
1–5 (proximal to 15) mm, densely short-hairy, septate-glandular.
2–10 (proximal to 30) mm, predominantly short-hairy, sometimes velutinous, not or sparsely, sometimes moderately, septate-glandular.
opening widely;
epicalyx bractlets ± elliptic, 4–6(–8) × 1–2 mm;
sepals spreading, (5–)7–10 mm, apex obtuse to acute, apiculate;
petals overlapping, spreading, cream-white to yellowish, broadly elliptic, (5–)7–9 × (5–)6–8 mm, ± equal to or slightly longer than sepals;
filaments 1.5–2.5 mm, anthers 0.7–1 mm;
styles thickened, 1 mm.
opening widely;
epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm;
sepals spreading, 3–8(–9) mm, apex usually acute, sometimes obtuse and apiculate;
petals ± overlapping, spreading, cream-white to bright yellow, broadly obovate, (3–)4–8(–10) × 3–8 mm, usually longer than, rarely equal to, sepals;
filaments 1–3(–3.5) mm, anthers 0.6–1 mm;
styles thickened, 1 mm.
light brown, 1 mm.
light brown, 1 mm.
= 14.
Drymocallis arguta
Drymocallis lactea
Drymocallis arguta is the only species of the genus occurring east of the Rocky Mountains, except for D. fissa in immediately adjacent prairies and the Black Hills of South Dakota. It is primarily a species of intact prairies and pasturelands throughout the Great Plains, extending as scattered populations in equivalent habitats throughout the northeastern states and adjacent provinces. Some of these states consider it a species of conservation concern (as Potentilla arguta, tall cinquefoil), and the sparsely hairy nature of some of these populations suggests that further taxonomic attention might be warranted. Drymocallis arguta occurs also in the Colorado Front Range, generally at elevations higher than D. fissa, with which it sometimes intergrades morphologically. It intergrades also with D. convallaria but tends to be larger, coarser, and more densely hairy with more acute leaflets.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Varieties 2 (2 in the flora).
Drymocallis lactea has had a well-established identity as Potentilla glandulosa var. (or subsp.) nevadensis; however, the epithet lactea has priority at species rank and avoids possible confusion with P. nevadensis Boissier. D. D. Keck (in J. Clausen et al. 1940) based his concept of P. glandulosa subsp. nevadensis on white-petaled plants with narrow inflorescences, the common form in the central Sierra Nevada and south. His map and annotations, however, include populations with yellow petals and widely branched inflorescences from northern California and adjacent Oregon. The two extremes merge in the northern Sierra Nevada, and populations in the North Coast Ranges of California also are problematic; the extremes are accordingly treated here as varieties of a single species. The inclusion by Keck of Washington in the species range might have been based on specimens here assigned to P. pseudorupestris.
Drymocallis lactea can be distinguished from sympatric species by its relatively large, overlapping petals, non-leafy inflorescences, and singly toothed leaflets. Pedicels and distal stems are usually densely short-hairy with relatively few or no glandular hairs (in contrast to D. pseudorupestris), and septate glands are usually absent from stem bases (in contrast to D. hansenii). The distinction between D. lactea and D. pseudorupestris breaks down at the margin of the species range in Nevada and Oregon.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Inflorescences narrow, branch angles 10–20°; petals cream-white to pale yellowish; pedicels and stems not or sparsely septate-glandular. | var. lactea |
1. Inflorescences wide, branch angles mostly 20–40°; petals usually pale to bright yellow; pedicels and stems sometimes moderately septate-glandular. | var. austiniae |
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