Drymocallis arguta |
Drymocallis convallaria |
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cordilleran drymocallis, sticky cinquefoil, tall cinquefoil, tall drymocallis or wood beauty, tall wood-beauty |
cordilleran drymocallis or wood beauty, cordilleran woodbeauty, cream cinquefoil, sharp tooth cinquefoil, valley cinquefoil, white cinquefoil |
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Caudex branches | short. |
short. |
Stems | usually solitary, sometimes tufted, (1–)3–10 dm; base 3–6 mm diam., ± densely, sometimes sparsely, septate-glandular. |
usually solitary, sometimes tufted, 4–9(–10) dm; base 2–5 mm diam., ± densely septate-glandular. |
Leaves | usually moderately to densely hairy; basal (6–)12–40 cm, leaflet pairs (3–)4–5; terminal leaflet broadly elliptic-obovate, (2–)4–10 × (1–)2–4.5 cm, teeth mostly double, 15–30+ per side, apex obtuse to acute; cauline 2–3, relatively well developed, leaflet pairs 2–4. |
sparsely to moderately, rarely densely, hairy; basal (10–)15–25(–30) cm, leaflet pairs (2–)3–4; terminal leaflet usually obovate, sometimes elliptic, 3–5(–6) × 1.5–4.5 cm, teeth double, 9–16 per side, apex rounded-obtuse; cauline (1–)2–4, well developed at least proximally, leaflet pairs usually 3. |
Inflorescences | 10–40-flowered, not leafy, congested, sometimes more elongate in fruit, 1/10–1/5(–1/3) of stem, narrow, branch angles 5–20°. |
(5–)10–40-flowered, not leafy, congested to open, (1/8–)1/6–1/3(–1/2) of stem, narrow, branch angles 5–20°. |
Pedicels | 1–5 (proximal to 15) mm, densely short-hairy, septate-glandular. |
1–5 (proximal to 25) mm, sparsely to ± densely short-hairy, predominantly septate-glandular. |
Flowers | opening widely; epicalyx bractlets ± elliptic, 4–6(–8) × 1–2 mm; sepals spreading, (5–)7–10 mm, apex obtuse to acute, apiculate; petals overlapping, spreading, cream-white to yellowish, broadly elliptic, (5–)7–9 × (5–)6–8 mm, ± equal to or slightly longer than sepals; filaments 1.5–2.5 mm, anthers 0.7–1 mm; styles thickened, 1 mm. |
opening ± widely; epicalyx bractlets narrowly elliptic, (2–)4–6 × 0.7–1.5 mm; sepals ± spreading, (4–)5–8(–10) mm, apex acute; petals not or scarcely overlapping, ± spreading, cream-white to pale yellow, obovate-elliptic, sometimes broadly so, (3–)5–8 × (2–)4–6 mm, ± equal to sepals; filaments 1.5–2.5 mm, anthers 0.7–1 mm; styles thickened, 0.8 mm. |
Achenes | light brown, 1 mm. |
light brown, 1 mm. |
2n | = 14. |
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Drymocallis arguta |
Drymocallis convallaria |
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Phenology | Flowering May–Aug. | Flowering (May–)Jun–Aug. |
Habitat | Prairies, abandoned pastures, open woodlands, shallow soil and grassy openings on ridges, slopes, bluffs, and rocky barrens | Dry to seasonally moist meadows, open forests, sagebrush and grassy rocky slopes |
Elevation | 10–2300 m (0–7500 ft) | 100–3000 m (300–9800 ft) |
Distribution |
AR; CO; CT; IA; IL; IN; KS; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NY; OH; OK; PA; SD; TN; VA; VT; WI; WV; WY; AB; MB; NB; NT; ON; QC; SK; YT
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AK; AZ; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; YT
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Discussion | Drymocallis arguta is the only species of the genus occurring east of the Rocky Mountains, except for D. fissa in immediately adjacent prairies and the Black Hills of South Dakota. It is primarily a species of intact prairies and pasturelands throughout the Great Plains, extending as scattered populations in equivalent habitats throughout the northeastern states and adjacent provinces. Some of these states consider it a species of conservation concern (as Potentilla arguta, tall cinquefoil), and the sparsely hairy nature of some of these populations suggests that further taxonomic attention might be warranted. Drymocallis arguta occurs also in the Colorado Front Range, generally at elevations higher than D. fissa, with which it sometimes intergrades morphologically. It intergrades also with D. convallaria but tends to be larger, coarser, and more densely hairy with more acute leaflets. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Drymocallis convallaria is widespread in western North America, occurring mostly west of the Continental Divide from south-central Alaska to Arizona. It differs from most sympatric species in usually having a single thick stem, a narrow inflorescence, and cream-white to pale yellow petals, at most only slightly longer than the sepals. Some collections from southern Idaho and northern Nevada that have very small petals in the size range of Drymocallis micropetala are included here on the basis of glandular pedicels. Excluded here are comparably small-petaled plants from the Warner Mountains of California and Oregon that have the aspect of D. convallaria but the blunter, redder achenes of D. glandulosa. Populations in northern Idaho and adjacent Washington that approach D. arguta in size and vestiture are provisionally treated as D. convallaria on the basis of leaflet shape. Specimens from New Mexico previously assigned to this taxon have largely been redetermined to D. arguta in the strict sense. See also discussions of Potentilla fissa var. major under 1. D. fissa, and of D. valida under the genus discussion. The illegitimate names P. glutinosa Nuttall ex Rydberg and D. glutinosa Rydberg have been applied most often to D. convallaria. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 285. | FNA vol. 9, p. 286. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Potentilla arguta, D. agrimonoides | Potentilla convallaria, D. arguta subsp. convallaria, P. arguta subsp. convallaria, P. arguta var. convallaria |
Name authority | (Pursh) Rydberg: Monogr. N. Amer. Potentilleae, 192. (1898) | (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 193. (1898) |
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