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Fendler's drymary

Habit Herbs [small shrubs], winter annual, annual, or perennial; taprooted, not rhizomatous.
Stems

prostrate to erect, simple or branched.

Leaves

opposite or sometimes appearing whorled, bases connate or not, sometimes petiolate or often sessile, stipulate;

stipules ovate or deltate to lanceolate or bristlelike, scarious;

blade subulate or subtriangular to linear and threadlike or spatulate to ovate or orbiculate, seldom succulent.

Inflorescences

terminal or axillary cymes, or flowers solitary;

bracts scarious or absent;

involucel bracteoles absent.

Pedicels

present or flowers sessile.

Flowers

bisexual or rarely unisexual;

perianth and androecium hypogynous or perigynous;

hypanthium absent or dish- or cup-shaped;

sepals 5, distinct or sometimes connate proximally, hooded (Drymaria, Polycarpon) or not, awned (Polycarpon) or not;

petals absent or (3–)5, blade clawed (Drymaria) or not, auricles absent, coronal appendages absent, blade apex entire, erose, or 2(–4)-fid, sometimes emarginate;

stamens (1–)3–5(–10), in 1 or 2 whorls, usually arising from base of ovary or from rim of hypanthium (Spergula, Spergularia);

staminodes absent;

ovary 1-locular;

styles 1 or 3, occasionally 2 or 5, distinct or sometimes connate proximally;

stigmas 3, occasionally 2 or 5.

Fruits

capsules, opening by 3 or 5, occasionally 4 valves;

carpophore sometimes present.

Seeds

3–150+, whitish or tan to often brown or black, ± triangular, pyriform, or reniform to circular, subglobose or laterally compressed to angular (Polycarpon);

embryo peripheral, curved or rarely annular to spirally curved (Spergula).

x

= [7], 8, 9, (11), 12.

Drymaria glandulosa

Caryophyllaceae subfam. polycarpoideae

Distribution
from FNA
AZ; NM; Mexico
[WildflowerSearch map]
[BONAP county map]
w North America (including Mexico); w South America; Europe (Mediterranean region); Asia (Mediterranean region); Africa (Mediterranean and tropical regions); less diverse in temperate areas
Discussion

Varieties 2 (1 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 16, species ca. 210 (7 genera, 28 species in the flora).

More commonly recognized as a tribe, Polycarpoideae is characterized by the presence of stipules, well-developed petals, and capsules. It is a relatively small group, with about four-fifths of the species in Drymaria, Polycarpaea, and Spergularia. Although clearly stipulate, Spergularia clusters with members of Alsinoideae in recent molecular studies (M. Nepokroeff et al. 2002; R. D. Smissen et al. 2002).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 12. FNA vol. 5, p. 9. Authors: Richard K. Rabeler, Ronald L. Hartman.
Parent taxa Caryophyllaceae > subfam. Polycarpoideae > Drymaria Caryophyllaceae
Sibling taxa
D. cordata, D. depressa, D. effusa, D. laxiflora, D. leptophylla, D. molluginea, D. pachyphylla, D. viscosa
Subordinate taxa
D. glandulosa var. glandulosa
Name authority Bartling: in C. B. Presl, Reliq. Haenk. 2: 9. (1831) Tanfani: in F. Parlatore, Fl. Ital. 9: 623. (1892)
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