Drummondia prorepens |
Orthotrichaceae |
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Drummond moss |
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Habit | Plants olive green, dark green, or yellow-brown. | Plants small to large, in tufts, cushions, or mats, dark green, reddish brown, or olive brown, dull. | ||||||||||||||||||||||||||||||||
Stem(s) | leaves channeled, concave, 1.1–1.8 mm; apex sometimes incurved; costa broadly channeled; alar cells yellow, ± inflated at marginal insertion; basal laminal cells 6–10 µm. Perichaetial leaves closely clasping seta, broadly lanceolate, to 2 mm, apex acute or acuminate, basal laminal cells rectangular, yellowish green. |
erect-ascending and often 2-fid, or creeping, branches many, erect or ascending, simple or 2-fid; outer cell walls thick, central strand absent, inner cells uniform, walls thin. |
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Leaves | erect-appressed, crowded, spirally twisted, flexuose, crisped, or contorted when dry, erect-spreading to squarrose-recurved when moist, ovate-lanceolate, ligulate-oblong, or lanceolate-linear, ± channeled; base not decurrent (decurrent in Zygodon); margins usually plane to revolute, rarely involute to erect, entire or sometimes denticulate near apex; apex rounded-obtuse, acute, occasionally acuminate, apiculate, or awned; costa strong, ending near apex; alar cells rarely differentiated; basal laminal cells rectangular, elongate-linear, quadrate, rounded, or elliptic; distal cells rounded-hexagonal, rarely rectangular, usually small, papillae 1–4(–6) per cell, conic or 2-fid, rarely smooth or mammillose, walls often incrassate. |
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Seta | dextrorse. |
erect, dextrorse or sinistrorse, smooth or rarely rough. |
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Sexual condition | usually gonioautoicous or dioicous, rarely pseudautoicous or cladautoicous; perigonia terminal or lateral, sometimes occurring on dwarf male plants, budlike, large; perichaetia terminal, further branching occurring by innovations, perichaetial leaves sometimes larger than stem leaves. |
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Capsule | 0.9–1.3 mm, sharply contracted to seta, neck absent; exostome teeth 3–6 cells high, rudimentary, truncate. |
erect, immersed, emergent, or exserted, ovate to cylindric-fusiform, symmetric, smooth or 8-ribbed, rarely 16-ribbed, sometimes constricted below mouth; exothecial cells rectangular to elliptic, often differentiated into bands; stomata superficial or immersed, usually below mid capsule, well developed; annulus poorly developed or absent; operculum convex to conic, rostrate; peristome double, single, rudimentary or rarely absent; prostome sometimes present, usually fragmentary; exostome teeth 16, usually connate in 8 pairs, erect, recurved, or reflexed, lanceolate, thick, densely papillose or striate; endostome segments 8 or 16 when present, hyaline, thin, linear-lanceolate, alternating with exostome teeth or connate to form rudimentary membrane, cilia absent. |
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Calyptra | mitrate or rarely cucullate, usually large and conspicuous, hairy or naked, plicate or not. |
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Spores | rounded-quadrate to elliptic-rectangular, 70–100 µm in longest dimension, almost smooth, brown. |
isosporous or anisosporous. |
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Specialized | asexual reproduction occasional, by gemmae. |
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Drummondia prorepens |
Orthotrichaceae |
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Habitat | Trunks and branches of deciduous trees, dry, upland forests, cedar glades, conifers, logs | |||||||||||||||||||||||||||||||||
Elevation | low to moderate elevations | |||||||||||||||||||||||||||||||||
Distribution |
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; SC; TN; VA; VT; WV; NB; NS; ON; QC
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Nearly worldwide |
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Discussion | Drummondia prorepens is characterized by long, creeping stems with many erect branches bearing terminal sporophytes. This habit, combined with the cucullate calyptrae, the smooth, ovate capsules with 16 truncate exostome teeth, the lack of stomata, and the undifferentiated basal laminal cells, is absolutely diagnostic. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 22, species ca. 380 (9 genera, 71 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 28, p. 41. | FNA vol. 28, p. 37. | ||||||||||||||||||||||||||||||||
Parent taxa | Orthotrichaceae > Drummondia | |||||||||||||||||||||||||||||||||
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Synonyms | Gymnostomum prorepens, D. canadensis, D. clavellata, D. clavellata var. canadensis | |||||||||||||||||||||||||||||||||
Name authority | (Hedwig) E. Britton: Mem. Torrey Bot. Club 4: 180. (1894) | Schimper | ||||||||||||||||||||||||||||||||
Web links |