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Siberian whitlowgrass

Macoun's Draba, Macoun's whitlow-grass

Habit Perennials; (stoloniferous); caudex branched (sparsely covered with persistent petiole remains, branches slender, creeping); scapose. Perennials; (cespitose); caudex simple or branched (with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose.
Stems

(subdecumbent when sterile), unbranched, 0.5–2.5 dm, sparsely to densely hispid, trichomes malpighiaceous, (flowering scapes sparsely pilose proximally with simple trichomes, often glabrous on distal 1/2).

unbranched, (0.06–)0.1–0.4(–0.6) dm, usually pubescent, rarely glabrous, trichomes simple and stalked, 2–4-rayed, 0.15–0.9 mm.

Basal leaves

rosulate;

petiolate;

blade oblong-lanceolate or oblanceolate, 0.4–2.2 cm × 1–5(–10) mm, margins entire, surfaces often pubescent, sometimes glabrous (except margins), with appressed, malpighiaceous trichomes (these sometimes with 1 or 2 shorter, lateral branches, appearing 3-fid or cross-shaped).

rosulate;

petiole (obscure), margin ciliate, (trichomes simple, 0.3–1 mm);

blade oblanceolate to obovate, (0.4–)0.6–1(–1.5) cm × (1–)2–4 mm, margins entire, surfaces usually pubescent abaxially, rarely glabrous, with short-stalked, 2–5-rayed trichomes, 0.1–0.5 mm, adaxially glabrous or pubescent with simple and short-stalked, 2-rayed trichomes.

Cauline leaves

0.

0.

Racemes

7–20-flowered, ebracteate, considerably elongated in fruit;

rachis not flexuous (straight), glabrous.

3–10(–13)-flowered, ebracteate, (subumbellate), slightly elongated in fruit;

rachis not flexuous, usually pubescent as stem, rarely glabrous.

Flowers

sepals (erect), oblong or ovate, 2–2.7 mm (lateral pair subsaccate basally), glabrous or abaxially sparsely pilose, (trichomes simple);

petals yellow, narrowly obovate, 4–6 × 2–3 mm, (apex emarginate);

anthers ovate-subcordate, 0.4–0.5 mm.

sepals ovate, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple and short-stalked, 2- or 3-rayed);

petals pale yellow, spatulate, 2.7–4 × 1–2 mm;

anthers ovate, 0.2–0.3 mm.

Fruiting pedicels

divaricate, straight or curved, (often filiform), 5–18(–23) mm, glabrous.

horizontal to divaricate (not expanded basally), curved upward, 1.5–4.5(–6) mm, abaxially pubescent as stem, adaxially usually glabrous, rarely throughout.

Fruits

usually oblong to elliptic, rarely sublinear, plane, flattened, 4–8 × 1.5–2.2 mm;

valves glabrous, (obscurely veined);

ovules 24–30 per ovary;

style 0.5–1 mm.

subglobose to ovoid or ellipsoid, plane, inflated at least basally, 4–8 × 2–4.5 mm;

valves glabrous;

ovules 10–14(–16) per ovary;

style 0.1–0.5 mm.

Seeds

(brown), ovoid, 0.9–1.2 × 0.5–0.6 mm.

ovoid, 1–1.2 × 0.6–0.8 mm.

2n

= 16.

= 64.

Draba sibirica

Draba macounii

Phenology Flowering May–Jul. Flowering Jun–Aug.
Habitat Wet places on rocky slopes Rock outcrops, talus, tundra
Elevation 0-300 m (0-1000 ft) 700-2800 m (2300-9200 ft)
Distribution
from USDA
Greenland; Europe (Russia); Asia (Caucasus, Iran, Russian Far East, Siberia, Turkey, central republics)
[BONAP county map]
from FNA
AK; MT; AB; BC; YT
[BONAP county map]
Discussion

Draba sibirica is one of three species of the genus in North America exhibiting malpighiaceous trichomes. It is easily distinguished from others (D. malpighiacea and some plants of D. spectabilis) by a complete absence of cauline leaves. It is found in the flora area in northeastern Greenland.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba macounii is often confused with the circumpolar decaploid (2n = 80) D. alpina. Plants of the former have scapes (0.6–)1–4(–6) cm, fruiting pedicels divaricate to horizontal, gently curved, and 1.5–4.5(–6) mm, and petals pale yellow, spatulate, and 2.7–4 × 1–2 mm. By contrast, D. alpina has scapes (3–)5–17(–28) cm, fruiting pedicels divaricate-ascending to ascending, often straight, and (3–)4–14(–30) mm, and petals bright yellow, obovate, and 3.5–5 × 1.7–2.5 mm. R. C. Rollins (1993) indicated that the species occurs in Colorado, but we have not seen any material from that state.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 337. FNA vol. 7, p. 318.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms Lepidium sibiricum, D. gmelinii, D. repens, D. sibirica subsp. arctica
Name authority (Pallas) Thellung: Neue Denkschr. Schweiz. Naturf. Ges. 41: 318. (1907) O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 97. (1927)
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