Draba santaquinensis
Draba cinerea
gray-leaf Draba, gray-leaf whitlow-grass
usually branched, 1.1–3.4 dm, hirsute proximally, trichomes mostly simple and 2-rayed, 0.2–1.5 mm, pubescent distally, trichomes mostly 2- or 3-rayed, 0.2–1 mm.
unbranched, (0.3–)0.5–1.6(–2.7) dm, pubescent throughout, trichomes 4–10-rayed, 0.1–0.3 mm, (often some rays branched, simple trichomes sparse, to 0.6 mm).
rosulate;
petiole (obscure), ciliate, (trichomes simple and 2-rayed);
blade obovate to oblanceolate, 1.6–2.5 cm × 5–10 mm, margins usually entire, rarely denticulate, surfaces pubescent, abaxially with stalked, mostly cruciform trichomes, 0.5–1 mm, adaxially with simple and stalked, 2–4-rayed ones.
rosulate;
petiolate;
petiole base and proximalmost margin ciliate, (trichomes simple, 0.2–0.8 mm);
blade oblanceolate to narrowly obovate or linear-lanceolate, 0.4–1.5 cm × 1–5 mm, margins usually entire, rarely with 1 tooth on each side, surfaces densely pubescent with minutely stalked, stellate, 8–12-rayed, trichomes 0.1–0.25 mm, (midvein obscure abaxially, apex trichomes simple).
usually 1–5;
sessile or subsessile;
blade oblanceolate to oblong, margins often denticulate, (ciliate proximally, with 2- or 3-rayed trichomes).
0–3(–5);
sessile;
blade ovate or oblong to lanceolate, margins entire, surfaces pubescent as basal.
9–25-flowered, ebracteate, elongated in fruit;
rachis not flexuous, pubescent, trichomes 2- or 3-rayed.
(3–)5–17(–24)-flowered, ebracteate, elongated in fruit;
rachis not flexuous, pubescent as stem.
sepals oblong-obovate, 2–3.5 mm, pubescent, (trichomes short- to long-stalked, 2–4-rayed, 0.2–0.8 mm);
petals yellow (often fading whitish), oblanceolate, 3–5 × 1–1.5 mm;
anthers ovate, 0.4–0.5 mm.
sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and short-stalked, 2- or 3-rayed);
petals white, spatulate to obovate, 3.5–4.5 × 1.5–2 mm;
anthers ovate, 0.3–0.4 mm.
horizontal to divaricate-ascending, usually straight, rarely curved upward, 7–10 mm, pubescent, trichomes 2- or 3-rayed.
divaricate-ascending or ascending, straight, (3–)4–7(–9) mm, pubescent as stem.
narrowly elliptic to slightly falcate, plane, flattened, (7–)10–16 × (2–)2.5–3.7 mm;
valves pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.5 mm;
ovules 14–22 per ovary;
style 0.9–1.2 mm.
oblong to elliptic, plane, slightly flattened, 5–8 × 2–3 mm;
valves pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.3 mm, (some rays branched);
ovules 20–36(–42) per ovary;
style (0.1–)0.2–1 mm.
oblong, 1.1–1.5 × 0.7–0.9 mm.
ovoid, 0.6–0.8 × 0.4–0.6 mm.
= 20.
= 48.
Draba santaquinensis
Draba cinerea
Although Draba santaquinensis was included within D. brachystylis by previous authors, I. A. Al-Shehbaz and M. D. Windham (2007) have shown that it is distinct both morphologically and chromosomally. It is currently known only from Utah County (American Fork, Provo, and Santaquin canyons) in north-central Utah.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Draba cinerea is a polymorphic species in which tetraploid, hexaploid, heptaploid, and octoploid chromosome counts (all based on x = 8) have been reported (e.g., R. C. Rollins 1993). Although we have not seen the type collection, we tentatively consider the taxon to be hexaploid, which is the predominant ploidy level throughout the species range, including North America, and also from its type region in Siberia.
Hexaploid Draba cinerea is closely related to decaploid (2n = 80) D. arctica, from which it is distinguished with difficulty. In general, D. cinerea has 1–3(–5)-leaved stems, basal leaf blades usually not terminated with simple trichomes and with obscure midveins abaxially, petals 3.5–4.5 mm, and seeds 0.6–0.8 × 0.4–0.6 mm. By contrast, D. arctica has leafless or 1(–3)-leaved stems, basal leaf blades terminated with simple or 2-rayed trichomes and with midveins distinct abaxially, petals 3.5–6 mm, and seeds (0.8–)0.9–1.1 × (0.6–)0.7–0.8 mm.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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