Draba santaquinensis |
Draba chamissonis |
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Cape Thompson Draba |
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Habit | Biennials or perennials; (short-lived); caudex simple (poorly developed, without persistent leaf bases); not scapose. | Perennials; (cespitose, sometimes forming mats); caudex branched (covered with persistent leaf bases); not scapose. |
Stems | usually branched, 1.1–3.4 dm, hirsute proximally, trichomes mostly simple and 2-rayed, 0.2–1.5 mm, pubescent distally, trichomes mostly 2- or 3-rayed, 0.2–1 mm. |
unbranched, 0.4–1.2 dm, pubescent throughout, trichomes minutely stalked, stellate, 8–12-rayed, (non-crisped), 0.03–0.3 mm, sometimes with simple and 2-rayed ones, to 0.3 mm. |
Basal leaves | rosulate; petiole (obscure), ciliate, (trichomes simple and 2-rayed); blade obovate to oblanceolate, 1.6–2.5 cm × 5–10 mm, margins usually entire, rarely denticulate, surfaces pubescent, abaxially with stalked, mostly cruciform trichomes, 0.5–1 mm, adaxially with simple and stalked, 2–4-rayed ones. |
rosulate; petiole (obsolete), margin usually not ciliate, (trichomes, when present, simple and 2-rayed, 0.2–0.35 mm); blade oblanceolate to obovate, 0.4–1 cm × 1–3 mm, margins denticulate or entire, surfaces pubescent with short-stalked, stellate, (non-crisped), 8–12-rayed trichomes, 0.05–0.2 mm, (midvein obscure abaxially). |
Cauline leaves | usually 1–5; sessile or subsessile; blade oblanceolate to oblong, margins often denticulate, (ciliate proximally, with 2- or 3-rayed trichomes). |
(1 or) 2–7; sessile; blade broadly ovate, margins usually denticulate, rarely entire. |
Racemes | 9–25-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent, trichomes 2- or 3-rayed. |
4–10-flowered, proximalmost 1–5 flowers bracteate, elongated; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong-obovate, 2–3.5 mm, pubescent, (trichomes short- to long-stalked, 2–4-rayed, 0.2–0.8 mm); petals yellow (often fading whitish), oblanceolate, 3–5 × 1–1.5 mm; anthers ovate, 0.4–0.5 mm. |
sepals ovate, 1.5–2.2 mm, pubescent, (trichomes simple and short-stalked, 2–5-rayed); petals white, obovate, 2–3.5 × 1.5–2 mm; anthers ovate, 0.2–0.3 mm. |
Fruiting pedicels | horizontal to divaricate-ascending, usually straight, rarely curved upward, 7–10 mm, pubescent, trichomes 2- or 3-rayed. |
ascending, straight, (4–)6–13(–20) mm, pubescent as stem. |
Fruits | narrowly elliptic to slightly falcate, plane, flattened, (7–)10–16 × (2–)2.5–3.7 mm; valves pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.5 mm; ovules 14–22 per ovary; style 0.9–1.2 mm. |
linear to narrowly lanceolate, slightly twisted or plane, flattened, 5–11 × 1.1–1.6(–2) mm; valves glabrous; ovules 16–20 per ovary; style 0.1–0.3 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–0.9 mm. |
ovoid, 0.6–0.9 × 0.3–0.5 mm. |
2n | = 20. |
= 16. |
Draba santaquinensis |
Draba chamissonis |
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Phenology | Flowering late Apr-early Jun. | Flowering Jun–Jul. |
Habitat | Limestone outcrops and rocky slopes in mixed conifer communities | Rock cliffs, bluffs, wind-swept tundra |
Elevation | 1800-2400 m (5900-7900 ft) | 0-500 m (0-1600 ft) |
Distribution |
UT |
AK; e Asia (Russian Far East, n Siberia) |
Discussion | Although Draba santaquinensis was included within D. brachystylis by previous authors, I. A. Al-Shehbaz and M. D. Windham (2007) have shown that it is distinct both morphologically and chromosomally. It is currently known only from Utah County (American Fork, Provo, and Santaquin canyons) in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
C. L. Hitchcock (1941) treated Draba chamissonis as a variety of D. nivalis and listed it from British Columbia; R. C. Rollins (1993) did not mention the taxon at any rank. The limited North American material of D. chamissonis that we examined is all from Alaska. The species strongly resembles D. nivalis, from which it is distinguished by having (1 or) 2–7 broadly ovate cauline leaves, proximalmost fruiting pedicels (4–)6–13(–20) mm, petals 1.5–2 mm wide, and fruits 1.1–1.6(–2) mm wide. By contrast, D. nivalis has one ovate or oblong cauline leaf, or none, proximalmost fruiting pedicels 1–4.5(–8) mm, petals 0.8–1.4 mm wide, and fruits 1.5–2.2 mm wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 334. | FNA vol. 7, p. 300. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. frigida var. kamtschatica, D. kamtschatica, D. lonchocarpa subsp. kamtschatica, D. nivalis var. kamtschatica | |
Name authority | Windham & Allphin: Harvard Pap. Bot. 12: 410. (2007) | G. Don: Gen. Hist. 1: 184. (1831) |
Web links |