Draba ruaxes |
Draba zionensis |
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coast mountain Draba, coast mountain whitlow-grass, coast mountain whitlow-grass Draba, Rainier Draba |
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Habit | Perennials; (cespitose, forming tufts); caudex branched (with persistent leaf bases, branches some terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, 0.2–0.6(–0.8) dm, often pubescent throughout, sometimes glabrate distally, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.4 mm. |
unbranched, 0.5–1.5(–1.8) dm, usually sparsely pubescent proximally, usually glabrous distally, rarely throughout, trichomes simple, 0.3–0.8 mm, often with smaller, 2–4-rayed ones. |
Basal leaves | rosulate; subsessile; petiole base and margin ciliate, (trichomes simple, 0.4–1.4 mm); blade oblanceolate to obovate or suborbicular, 0.3–1 cm × 2–4.5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–10-rayed, stellate trichomes, 0.2–0.8 mm, adaxially with simple trichomes, 0.4–1 mm, sometimes with smaller, 2–4-rayed ones, (midvein obscure). |
rosulate; shortly petiolate; petiole base sometimes ciliate (margin not ciliate, trichomes simple and 2-rayed, 0.2–0.5 mm); blade spatulate to obovate, 0.7–3.5 cm × 3–10(–12) mm, margins usually entire, rarely obscurely dentate (near apex), surfaces sparsely pubescent with stalked, (2–)4 (or 5)-rayed trichomes, 0.05–0.5 mm, (midvein obscure). |
Cauline leaves | 0. |
0. |
Racemes | (2–)4–10(–14)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem or glabrous. |
14–36-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals ovate, 2–3 mm, pubescent, (trichomes simple with fewer, 2-rayed ones); petals bright yellow, obovate, 4–6 × 2–3.5 mm; anthers oblong, 0.4–0.5 mm. |
sepals ovate, 2–3 mm, glabrous or sparsely pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals orange-yellow (fading pale yellow), broadly spatulate to subovate, 5–6 × 1.8–2.8 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or slightly curved upward, 3–7(–9) mm, pubescent or glabrous, trichomes simple and 2–4-rayed. |
ascending to divaricate-ascending, straight, (not expanded basally), (4–)5–12(–15) mm, glabrous. |
Fruits | elliptic to lanceolate or ovate to suborbicular, plane, flattened, 4–8(–10) × 3–4.5 mm; valves puberulent, trichomes simple with fewer 2-rayed ones, 0.1–0.35 mm; ovules 12–16 per ovary; style 0.5–0.9(–1.1) mm. |
lanceolate to linear-lanceolate or ovate, plane (not curved), flattened, 6–12(–17) × 2–3 mm; valves glabrous; ovules 12–20 per ovary; style 0.4–1 mm. |
Seeds | oblong, 1.5–2 × 1–1.3 mm. |
oblong, 1–1.3 × 0.6–0.8 mm. |
2n | = 72. |
= 26. |
Draba ruaxes |
Draba zionensis |
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Phenology | Flowering Jun–Jul. | Flowering Mar–May. |
Habitat | Rock outcrops, talus slopes, ridges, alpine summits | Sandstone (rarely limestone) rock outcrops and sandy slopes in pinyon-juniper or pine communities |
Elevation | 500-2400 m (1600-7900 ft) | 1000-2500 m (3300-8200 ft) |
Distribution |
AK; WA; AB; BC; YT
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UT |
Discussion | C. L. Hitchcock (1941) treated Draba ruaxes as a variety of D. ventosa; as demonstrated by G. A. Mulligan (1971b), the two are quite distinct. Draba ruaxes is an outcrossing hexaploid with well-formed anthers and pollen, and abundant, simple trichomes on leaves, stems, sepals, and fruits. By contrast, D. ventosa is an apomictic triploid with abortive anthers and/or pollen, and no simple trichomes anywhere on the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
R. C. Rollins (1993) treated Draba zionensis as a variety of D. asprella, but its true relationships appear to lie with two other southern Utah endemics, D. sobolifera and D. subalpina. Draba zionensis is easily distinguished from D. subalpina by having orange-yellow (versus white) petals, and from D. asprella and D. sobolifera by its glabrous (versus pubescent) pedicels and stems distally. Nearly all populations of the species are found in and around Zion National Park in southwestern Utah (Iron, Kane, and Washington counties). A specimen supposedly from the Deep Creek Mountains (Juab County) may be mislabeled. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 334. | FNA vol. 7, p. 346. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. ventosa var. ruaxes | D. asprella var. zionensis |
Name authority | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 117. (1930) | C. L. Hitchcock: Revis. Drabas W. N. Amer., 49, plate 2, fig. 16. (1941) |
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