Draba ruaxes |
Draba pedicellata |
|
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coast mountain Draba, coast mountain whitlow-grass, coast mountain whitlow-grass Draba, Rainier Draba |
Cusick's Draba, rocky-slope Draba, rocky-slope whitlow-grass, stalk whitlow-cress |
|
Habit | Perennials; (cespitose, forming tufts); caudex branched (with persistent leaf bases, branches some terminating in sterile rosettes); scapose. | Perennials; (loosely cespitose); caudex branched (with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, 0.2–0.6(–0.8) dm, often pubescent throughout, sometimes glabrate distally, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.4 mm. |
unbranched, (0.15–)0.3–1.3(–1.6) dm, pubescent proximally, glabrous or nearly so distally, trichomes 2- or 3-rayed and simple, 0.1–1 mm. |
Basal leaves | rosulate; subsessile; petiole base and margin ciliate, (trichomes simple, 0.4–1.4 mm); blade oblanceolate to obovate or suborbicular, 0.3–1 cm × 2–4.5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–10-rayed, stellate trichomes, 0.2–0.8 mm, adaxially with simple trichomes, 0.4–1 mm, sometimes with smaller, 2–4-rayed ones, (midvein obscure). |
rosulate; subsessile; blade oblanceolate to obovate, 0.4–1.6(–2) cm × 2–4(–6) mm, margins entire, (not ciliate), surfaces pubescent with stalked, 2–4-rayed trichomes, 0.2–0.9 mm, rarely adaxially with simple trichomes, (midvein obscure abaxially). |
Cauline leaves | 0. |
0. |
Racemes | (2–)4–10(–14)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem or glabrous. |
3–20(–27)-flowered, ebracteate, slightly to considerably elongated in fruit; rachis straight or flexuous, usually glabrous, rarely pubescent, trichomes simple and 2–4-rayed. |
Flowers | sepals ovate, 2–3 mm, pubescent, (trichomes simple with fewer, 2-rayed ones); petals bright yellow, obovate, 4–6 × 2–3.5 mm; anthers oblong, 0.4–0.5 mm. |
sepals broadly ovate, 1.8–2.5 mm, glabrous or pubescent, (trichomes simple and branched); petals yellow, spatulate, 3.5–5.5 × 1.5–2.2 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or slightly curved upward, 3–7(–9) mm, pubescent or glabrous, trichomes simple and 2–4-rayed. |
divaricate to horizontal, straight or curved upward (expanded basally or not), 4–9(–13) mm, glabrous or pubescent, trichomes simple and 2–4-rayed. |
Fruits | elliptic to lanceolate or ovate to suborbicular, plane, flattened, 4–8(–10) × 3–4.5 mm; valves puberulent, trichomes simple with fewer 2-rayed ones, 0.1–0.35 mm; ovules 12–16 per ovary; style 0.5–0.9(–1.1) mm. |
lanceolate to ellipsoid or ovoid, plane (not curved), inflated or slightly flattened, (4.2–)5.3–10(–13) × 2.5–4.5(–5.5) mm; valves glabrous or pubescent, trichomes simple and short-stalked, 2-rayed, 0.05–0.3 mm; ovules 8–16 per ovary; style 0.3–1(–1.2) mm. |
Seeds | oblong, 1.5–2 × 1–1.3 mm. |
oblong, 1.2–1.6 × 0.7–0.9 mm. |
2n | = 72. |
= 20. |
Draba ruaxes |
Draba pedicellata |
|
Phenology | Flowering Jun–Jul. | Flowering May–Jul. |
Habitat | Rock outcrops, talus slopes, ridges, alpine summits | Rock outcrops, talus, steep gravel slopes in pinyon-juniper, mixed conifer, and subalpine meadow communities |
Elevation | 500-2400 m (1600-7900 ft) | 2300-3800 m (7500-12500 ft) |
Distribution |
AK; WA; AB; BC; YT
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NV; UT
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Discussion | C. L. Hitchcock (1941) treated Draba ruaxes as a variety of D. ventosa; as demonstrated by G. A. Mulligan (1971b), the two are quite distinct. Draba ruaxes is an outcrossing hexaploid with well-formed anthers and pollen, and abundant, simple trichomes on leaves, stems, sepals, and fruits. By contrast, D. ventosa is an apomictic triploid with abortive anthers and/or pollen, and no simple trichomes anywhere on the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba pedicellata is known from Elko, Eureka, Nye, and White Pine counties in northeastern Nevada and to Tooele County in northwestern Utah. It was considered a variety of D. cusickii by R. C. Rollins (1993); chromosome number, morphology, and geographic distribution suggest a closer relationship to D. sphaeroides. For a detailed discussion and comparison of these species, consult M. D. Windham (2004). N. H. Holmgren (2005b) divided D. pedicellata into two varieties: var. pedicellata, which is found on limestone throughout the species range, and var. wheelerensis, which is found on non-calcareous substrates in White Pine County. Although var. wheelerensis has slightly more spreading pedicels on somewhat more flexuous rachises, it appears to be little more than a dwarf alpine form of the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 334. | FNA vol. 7, p. 328. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. ventosa var. ruaxes | D. cusickii var. pedicellata, D. pedicellata var. wheelerensis |
Name authority | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 117. (1930) | (Rollins & R. A. Price) Windham: Madroño 50: 221. (2004) |
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