Draba ruaxes |
Draba incerta |
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coast mountain Draba, coast mountain whitlow-grass, coast mountain whitlow-grass Draba, Rainier Draba |
whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
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Habit | Perennials; (cespitose, forming tufts); caudex branched (with persistent leaf bases, branches some terminating in sterile rosettes); scapose. | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, 0.2–0.6(–0.8) dm, often pubescent throughout, sometimes glabrate distally, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.4 mm. |
unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
Basal leaves | rosulate; subsessile; petiole base and margin ciliate, (trichomes simple, 0.4–1.4 mm); blade oblanceolate to obovate or suborbicular, 0.3–1 cm × 2–4.5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–10-rayed, stellate trichomes, 0.2–0.8 mm, adaxially with simple trichomes, 0.4–1 mm, sometimes with smaller, 2–4-rayed ones, (midvein obscure). |
rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
Cauline leaves | 0. |
usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
Racemes | (2–)4–10(–14)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem or glabrous. |
3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
Flowers | sepals ovate, 2–3 mm, pubescent, (trichomes simple with fewer, 2-rayed ones); petals bright yellow, obovate, 4–6 × 2–3.5 mm; anthers oblong, 0.4–0.5 mm. |
sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or slightly curved upward, 3–7(–9) mm, pubescent or glabrous, trichomes simple and 2–4-rayed. |
ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
Fruits | elliptic to lanceolate or ovate to suborbicular, plane, flattened, 4–8(–10) × 3–4.5 mm; valves puberulent, trichomes simple with fewer 2-rayed ones, 0.1–0.35 mm; ovules 12–16 per ovary; style 0.5–0.9(–1.1) mm. |
broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
Seeds | oblong, 1.5–2 × 1–1.3 mm. |
oblong, 1.1–1.5 × 0.7–1 mm. |
2n | = 72. |
= 112. |
Draba ruaxes |
Draba incerta |
|
Phenology | Flowering Jun–Jul. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus slopes, ridges, alpine summits | Rock outcrops, talus, gravelly areas, tundra |
Elevation | 500-2400 m (1600-7900 ft) | 0-3300 m (0-10800 ft) |
Distribution |
AK; WA; AB; BC; YT
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AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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Discussion | C. L. Hitchcock (1941) treated Draba ruaxes as a variety of D. ventosa; as demonstrated by G. A. Mulligan (1971b), the two are quite distinct. Draba ruaxes is an outcrossing hexaploid with well-formed anthers and pollen, and abundant, simple trichomes on leaves, stems, sepals, and fruits. By contrast, D. ventosa is an apomictic triploid with abortive anthers and/or pollen, and no simple trichomes anywhere on the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 334. | FNA vol. 7, p. 313. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. ventosa var. ruaxes | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei |
Name authority | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 117. (1930) | Payson: Amer. J. Bot. 4: 261. (1917) |
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