Draba ruaxes |
Draba hitchcockii |
|
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coast mountain Draba, coast mountain whitlow-grass, coast mountain whitlow-grass Draba, Rainier Draba |
lost river Draba |
|
Habit | Perennials; (cespitose, forming tufts); caudex branched (with persistent leaf bases, branches some terminating in sterile rosettes); scapose. | Perennials; (densely cespitose); caudex branched (densely covered with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, 0.2–0.6(–0.8) dm, often pubescent throughout, sometimes glabrate distally, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.4 mm. |
unbranched, (0.1–)0.3–1(–1.3) dm, hirsute throughout, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.6 mm. |
Basal leaves | rosulate; subsessile; petiole base and margin ciliate, (trichomes simple, 0.4–1.4 mm); blade oblanceolate to obovate or suborbicular, 0.3–1 cm × 2–4.5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–10-rayed, stellate trichomes, 0.2–0.8 mm, adaxially with simple trichomes, 0.4–1 mm, sometimes with smaller, 2–4-rayed ones, (midvein obscure). |
(densely imbricate); rosulate; sessile; blade narrowly oblanceolate to oblong-linear, 0.3–1.2(–1.5) cm × 1–2 mm, margins entire, (ciliate, trichomes simple, 0.3–1.2 mm), surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.08–0.45 mm, (midvein obscure), adaxially glabrous proximally, sparsely pubescent distally with mostly simple trichomes. |
Cauline leaves | 0. |
0. |
Racemes | (2–)4–10(–14)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem or glabrous. |
4–15-flowered, ebracteate, elongated in fruit; rachis not flexuous, hirsute as stem. |
Flowers | sepals ovate, 2–3 mm, pubescent, (trichomes simple with fewer, 2-rayed ones); petals bright yellow, obovate, 4–6 × 2–3.5 mm; anthers oblong, 0.4–0.5 mm. |
sepals oblong, 2.5–3.5 mm, hirsute, (trichomes simple and stalked, 2–4-rayed); petals white, obovate, 5–6.5 × 2–3.5 mm; anthers ovate, 0.5–0.6 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or slightly curved upward, 3–7(–9) mm, pubescent or glabrous, trichomes simple and 2–4-rayed. |
ascending, straight, (2–)4–13(–18) mm, hirsute as stem. |
Fruits | elliptic to lanceolate or ovate to suborbicular, plane, flattened, 4–8(–10) × 3–4.5 mm; valves puberulent, trichomes simple with fewer 2-rayed ones, 0.1–0.35 mm; ovules 12–16 per ovary; style 0.5–0.9(–1.1) mm. |
ovate to broadly oblong or elliptic, plane, flattened, (3–)4–7(–10) × (2.5–)3.5–5 mm; valves pubescent, trichomes simple and short-stalked, 2(–4)-rayed, 0.06–0.5 mm; ovules 8–12 per ovary; style (0.8–)1–1.7(–2) mm. |
Seeds | oblong, 1.5–2 × 1–1.3 mm. |
oblong, 1.4–1.8 × 0.8–1 mm. |
2n | = 72. |
= 54. |
Draba ruaxes |
Draba hitchcockii |
|
Phenology | Flowering Jun–Jul. | Flowering May–Jun. |
Habitat | Rock outcrops, talus slopes, ridges, alpine summits | Limestone outcrops and gravelly soil |
Elevation | 500-2400 m (1600-7900 ft) | 1800-2200 m (5900-7200 ft) |
Distribution |
AK; WA; AB; BC; YT
|
ID |
Discussion | C. L. Hitchcock (1941) treated Draba ruaxes as a variety of D. ventosa; as demonstrated by G. A. Mulligan (1971b), the two are quite distinct. Draba ruaxes is an outcrossing hexaploid with well-formed anthers and pollen, and abundant, simple trichomes on leaves, stems, sepals, and fruits. By contrast, D. ventosa is an apomictic triploid with abortive anthers and/or pollen, and no simple trichomes anywhere on the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba hitchcockii is known from the Lost River Range in Butte and Custer counties. Based on morphological and chromosomal evidence, M. D. Windham (2004) suggested that it may be an allopolyploid resulting from hybridization between D. oreibata and D. paysonii. Draba hitchcockii is superficially similar to D. jaegeri, a taxon known from the Charleston Mountains of Clark County, Nevada. Both are cespitose perennials with relatively large, white flowers and a chromosome number (2n = 54) otherwise unknown in Draba (Windham). Features distinguishing these two taxa are provided in the discussion of 51. D. jaegeri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 334. | FNA vol. 7, p. 311. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. ventosa var. ruaxes | |
Name authority | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 117. (1930) | Rollins: J. Arnold Arbor. 64: 500. (1983) |
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