Draba ruaxes |
Draba burkei |
|
---|---|---|
coast mountain Draba, coast mountain whitlow-grass, coast mountain whitlow-grass Draba, Rainier Draba |
snowbasin Draba |
|
Habit | Perennials; (cespitose, forming tufts); caudex branched (with persistent leaf bases, branches some terminating in sterile rosettes); scapose. | Perennials; (cespitose, forming loose mats); caudex branched (with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, 0.2–0.6(–0.8) dm, often pubescent throughout, sometimes glabrate distally, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.4 mm. |
unbranched, 0.3–0.6(–0.9) dm, glabrous. |
Basal leaves | rosulate; subsessile; petiole base and margin ciliate, (trichomes simple, 0.4–1.4 mm); blade oblanceolate to obovate or suborbicular, 0.3–1 cm × 2–4.5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–10-rayed, stellate trichomes, 0.2–0.8 mm, adaxially with simple trichomes, 0.4–1 mm, sometimes with smaller, 2–4-rayed ones, (midvein obscure). |
subrosulate; subsessile; blade oblanceolate to obovate, 0.3–0.8(–1.3) cm × 1–2.5 mm, margins entire, (ciliate, trichomes simple, subsetiform, 0.25–0.8 mm), surfaces glabrous. |
Cauline leaves | 0. |
0. |
Racemes | (2–)4–10(–14)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem or glabrous. |
4–10-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals ovate, 2–3 mm, pubescent, (trichomes simple with fewer, 2-rayed ones); petals bright yellow, obovate, 4–6 × 2–3.5 mm; anthers oblong, 0.4–0.5 mm. |
sepals broadly ovate, 2–3.5 mm, glabrous or sparsely pubescent, (trichomes simple, 0.07–0.35 mm); petals yellow, oblanceolate, 4–6 × 1.5–2 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | ascending to divaricate-ascending, straight or slightly curved upward, 3–7(–9) mm, pubescent or glabrous, trichomes simple and 2–4-rayed. |
divaricate-ascending to ascending, straight, 4–9(–15) mm, glabrous. |
Fruits | elliptic to lanceolate or ovate to suborbicular, plane, flattened, 4–8(–10) × 3–4.5 mm; valves puberulent, trichomes simple with fewer 2-rayed ones, 0.1–0.35 mm; ovules 12–16 per ovary; style 0.5–0.9(–1.1) mm. |
ovate, plane, flattened, 3–5.5 × 2–3.2 mm; valves glabrous or puberulent, trichomes simple, 0.02–0.08 mm; ovules 4–10 per ovary; style 0.5–1.7 mm. |
Seeds | oblong, 1.5–2 × 1–1.3 mm. |
ovoid, 1–1.4 × 0.7–1 mm. |
2n | = 72. |
= 20. |
Draba ruaxes |
Draba burkei |
|
Phenology | Flowering Jun–Jul. | Flowering May–Jul. |
Habitat | Rock outcrops, talus slopes, ridges, alpine summits | Rocky ridges, steep talus slopes, rock outcrops and crevices |
Elevation | 500-2400 m (1600-7900 ft) | 1600-3000 m (5200-9800 ft) |
Distribution |
AK; WA; AB; BC; YT
|
UT |
Discussion | C. L. Hitchcock (1941) treated Draba ruaxes as a variety of D. ventosa; as demonstrated by G. A. Mulligan (1971b), the two are quite distinct. Draba ruaxes is an outcrossing hexaploid with well-formed anthers and pollen, and abundant, simple trichomes on leaves, stems, sepals, and fruits. By contrast, D. ventosa is an apomictic triploid with abortive anthers and/or pollen, and no simple trichomes anywhere on the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba burkei was treated by C. L. Hitchcock (1941) and R. C. Rollins (1993) as a variety of D. maguirei. Chromosome numbers (2n = 20 versus 2n = 32), plant morphology, and molecular data support the recognition of these taxa as independent species (M. D. Windham 2004). Draba burkei is easily distinguished from D. maguirei by having exclusively simple trichomes confined to leaf blade margins (versus mostly branched trichomes on margins and surfaces) and smaller seeds (1–1.4 × 0.7–1 versus 1.6–2 × 1–1.3 mm). It is known from Box Elder, Cache, Morgan, and Weber counties, where it approaches but does not overlap the range of D. maguirei. Draba burkei (as D. maguirei var. burkei) is in the Center for Plant Conservation’s National Collection of Endangered Plants and is listed in NatureServe as a plant of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 334. | FNA vol. 7, p. 298. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. ventosa var. ruaxes | D. maguirei var. burkei |
Name authority | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 117. (1930) | (C. L. Hitchcock) Windham & Beilstein: Madroño 50: 221. (2004) |
Web links |