Draba ramulosa |
Draba oreibata |
|
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Tushar Mountain Draba |
limestone Draba |
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Habit | Perennials; (loosely matted, grayish); caudex branched (with persistent leaf bases, branches creeping, sometimes terminating in sterile rosettes); not scapose. | Perennials; (cespitose); caudex branched (with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, 0.4–0.6 dm, densely pubescent throughout, trichomes dendritic, 3–6-rayed, (often crisped), 0.1–0.4 mm. |
unbranched, 0.2–0.9 dm, glabrous throughout. |
Basal leaves | (imbricate); not rosulate; sessile; blade obovate to oblanceolate, 0.4–1.1 cm × 2–3.2 mm, margins entire, (base and margins not ciliate), surfaces pubescent with stalked, 4–8-rayed trichomes, 0.1–0.5 mm, (sometimes 1 or more rays spurred), adaxially sometimes trichomes simple. |
rosulate; sessile; blade oblong to narrowly oblanceolate, 0.2–1 cm × 0.6–2(–4) mm, margins entire, (ciliate, trichomes simple, 0.1–0.5 mm, apex obtuse), surfaces glabrous. |
Cauline leaves | 2 or 3 (sometimes basal leaves spaced, flowering stem appearing to 8-leaved); sessile; blade oblong to ovate, margins entire, surfaces pubescent as basal. |
0. |
Racemes | 4–15-flowered, ebracteate or proximalmost 1 or 2 flowers bracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
3–8-flowered, ebracteate, considerably elongated in fruit; rachis slightly flexuous, glabrous. |
Flowers | sepals broadly ovate, 1.7–2.4 mm, pubescent, (trichomes short-stalked, 2–5-rayed); petals yellow, obovate to oblanceolate, 3–4.5 × 1.5–2.5 mm; anthers ovate, 0.4–0.5 mm. |
sepals oblong, 1.7–2.5 mm, glabrous; petals white, oblanceolate to obovate, 3.5–4.5 × 1.2–2 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | divaricate-ascending, usually straight, rarely curved upward, 3–6(–10) mm, pubescent, trichomes 3–6-rayed, (crisped, 0.1–0.4 mm), and, sometimes, simple. |
divaricate-ascending to ascending, straight, 4–13 mm, glabrous. |
Fruits | (not appressed to rachis), ovate to elliptic, plane, flattened, 4–6.5 × 2.5–4 mm; valves densely pubescent, trichomes simple and short-stalked, 2–5-rayed, 0.08–0.35 mm; ovules 6–12 per ovary; style (0.1–)0.3–0.7 mm. |
oblong to narrowly so, slightly twisted, flattened, 5–9 × 1.5–3 mm; valves glabrous; ovules 8–16 per ovary; style 0.3–0.9(–1.2) mm. |
Seeds | oblong, 1.4–1.8 × 0.8–1.2 mm. |
oblong, 0.9–1.3 × 0.6–0.8 mm. |
2n | = 32. |
|
Draba ramulosa |
Draba oreibata |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Rock outcrops, talus, gravelly soils | Limestone cliffs, talus |
Elevation | 3300-3600 m (10800-11800 ft) | 1800-2700 m (5900-8900 ft) |
Distribution |
UT |
ID |
Discussion | Of conservation concern. Molecular and chromosomal data (M. D. Windham, unpubl.) strongly suggest that Draba ramulosa is an allopolyploid species. It is thought to have originated through hybridization between D. sobolifera and a member of the white-flowered, euploid lineage of M. A. Beilstein and M. D. Windham (2003). It is easily distinguished from D. sobolifera by having pale yellow to whitish (versus bright yellow) petals, grayish (versus green) foliage, non-ciliate (versus ciliate) basal leaves pubescent with 4–8-rayed (versus 2–4-rayed) trichomes, often proximally bracteate (versus ebracteate) racemes, and flattened (versus inflated basally) fruits. Draba ramulosa is known from the Tushar Mountains in south-central Utah (Beaver and Piute counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
R. C. Rollins (1993) and N. H. Holmgren (2005b) divided Draba oreibata into two varieties: the Idaho endemic var. oreibata and the Nevada endemic var. serpentina. As indicated by I. A. Al-Shehbaz and M. D. Windham (2007), the two taxa are morphologically distinct, have different chromosome numbers, and are separated by over 480 kilometers. They are treated herein as separate species, and distinguished by characteristics discussed by Al-Shehbaz and Windham. Draba oreibata, in the strict sense, is known from Blaine, Butte, Clark, Custer, and Lemhi counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 332. | FNA vol. 7, p. 325. |
Parent taxa | ||
Sibling taxa | ||
Name authority | Rollins: Contr. Gray Herb. 214: 6. (1984) | J. F. Macbride & Payson: Amer. J. Bot. 4: 257. (1917) |
Web links |