Flora of North America species comparison

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Draba ramulosa

Draba novolympica

Tushar Mountain Draba

Mt. Olympic Draba, Olympic Draba, Olympic Mountains Draba, Payson's Draba, Payson's whitlow-grass, Trelease's Draba, Trelease's whitlow-grass
Habit Perennials; (loosely matted, grayish); caudex branched (with persistent leaf bases, branches creeping, sometimes terminating in sterile rosettes); not scapose. Perennials; (cespitose, densely pulvinate); caudex branched (covered with persistent leaves, branches creeping, sometimes terminating in sterile rosettes); scapose.
Stems

unbranched, 0.4–0.6 dm, densely pubescent throughout, trichomes dendritic, 3–6-rayed, (often crisped), 0.1–0.4 mm.

unbranched, 0.05–0.4 dm, densely pubescent throughout, trichomes simple, 0.4–0.8 mm, and stalked, 2–5-rayed, 0.1–0.5 mm.
Basal leaves

(imbricate);

not rosulate;

sessile;

blade obovate to oblanceolate, 0.4–1.1 cm × 2–3.2 mm, margins entire, (base and margins not ciliate), surfaces pubescent with stalked, 4–8-rayed trichomes, 0.1–0.5 mm, (sometimes 1 or more rays spurred), adaxially sometimes trichomes simple.

(densely imbricate); rosulate;

sessile;

blade oblong to linear-oblanceolate, 0.2–0.8 cm × 0.5–1.5 mm, margins entire, (ciliate, trichomes simple and spurred, 0.3–1.2 mm), surfaces densely pubescent, abaxially with stalked, 2–12-rayed stellate trichomes, 0.1–0.6 mm, adaxially with simple and 2-rayed ones, 0.3–0.8 mm.
Cauline leaves

2 or 3 (sometimes basal leaves spaced, flowering stem appearing to 8-leaved);

sessile;

blade oblong to ovate, margins entire, surfaces pubescent as basal.

0.
Racemes

4–15-flowered, ebracteate or proximalmost 1 or 2 flowers bracteate, slightly elongated in fruit;

rachis not flexuous, pubescent as stem.

2–12-flowered, ebracteate, slightly elongated in fruit;

rachis not flexuous, pubescent as stem.
Flowers

sepals broadly ovate, 1.7–2.4 mm, pubescent, (trichomes short-stalked, 2–5-rayed);

petals yellow, obovate to oblanceolate, 3–4.5 × 1.5–2.5 mm;

anthers ovate, 0.4–0.5 mm.

sepals oblong, 1.5–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed);

petals bright yellow, oblanceolate to spatulate, 2–3.5(–4) × 1.5–2 mm;

anthers oblong, 0.5–0.6 mm.
Fruiting pedicels

divaricate-ascending, usually straight, rarely curved upward, 3–6(–10) mm, pubescent, trichomes 3–6-rayed, (crisped, 0.1–0.4 mm), and, sometimes, simple.

divaricate-ascending, straight, 1–5 mm, pubescent, trichomes simple (0.3–0.9 mm) and stalked, 2–5-rayed (0.1–0.5 mm).
Fruits

(not appressed to rachis), ovate to elliptic, plane, flattened, 4–6.5 × 2.5–4 mm;

valves densely pubescent, trichomes simple and short-stalked, 2–5-rayed, 0.08–0.35 mm;

ovules 6–12 per ovary;

style (0.1–)0.3–0.7 mm.

often ovoid, plane, slightly inflated basally (symmetric), (2.5–)3–4(–5) × 1.5–3.5 mm;

valves densely pubescent, trichomes 2–6-rayed, 0.05–0.4 mm, occasionally some simple;

ovules 4–8(–12) per ovary;

style 0.2–0.6(–0.8) mm.
Seeds

oblong, 1.4–1.8 × 0.8–1.2 mm.

oblong, 1.2–1.8 × 0.8–1.1 mm.
2n

= 42.

Draba ramulosa

Draba novolympica
Phenology Flowering Jun–Aug. Flowering Jun–Aug.
Habitat Rock outcrops, talus, gravelly soils Alpine crests, open knolls, fellfields, talus, weathered shale, calcareous shale scree, rocky grounds and cliffs, subalpine conifer forests
Elevation 3300-3600 m (10800-11800 ft) 1500-3700 m (4900-12100 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Molecular and chromosomal data (M. D. Windham, unpubl.) strongly suggest that Draba ramulosa is an allopolyploid species. It is thought to have originated through hybridization between D. sobolifera and a member of the white-flowered, euploid lineage of M. A. Beilstein and M. D. Windham (2003). It is easily distinguished from D. sobolifera by having pale yellow to whitish (versus bright yellow) petals, grayish (versus green) foliage, non-ciliate (versus ciliate) basal leaves pubescent with 4–8-rayed (versus 2–4-rayed) trichomes, often proximally bracteate (versus ebracteate) racemes, and flattened (versus inflated basally) fruits. Draba ramulosa is known from the Tushar Mountains in south-central Utah (Beaver and Piute counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba novolympica is the same taxon that C. L. Hitchcock (1941) and R. C. Rollins (1993) called D. paysonii var. treleasei, and G. A. Mulligan (2002) called D. paysonii. The two are amply distinct and should be recognized as separate species. Draba novolympica is easily distinguished from D. paysonii by having fruit valves pubescent with 2–6-rayed (occasionally some simple) trichomes 0.05–0.4 mm, sepals 1.5–2.5 mm, petals 2–3.5(–4) × 1.5–2 mm, fruits (2.5–)3–4(–5) × 1.5–3.5 mm, styles 0.2–0.6(–0.8) mm, and ovules 1.2–1.8 × 0.8–1.1 mm. By contrast, D. paysonii has fruit valves pubescent with simple and 2-rayed (some 4- or 5-rayed) trichomes (0.2–)0.4–1 mm, sepals 2.8–3.5 mm, petals (4–)5–6 × (1.5–)2–3 mm, fruits (5–)6–9 × (3–)3.5–5 mm, styles (0.6–)0.8–1.2 mm, and ovules 1.7–2.2 × 1–1.4 mm.

Both R. C. Rollins (1993) and N. H. Holmgren (2005b) indicated that Draba novolympica (as D. paysonii var. treleasei) occurs in Alaska and Yukon, but we have not seen any material from there, and it is likely that their records were based on misidentified plants. Previous reports of D. paysonii from Canada (e.g., G. A. Mulligan 1971b) pertain instead to D. novolympica.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 7, p. 332. FNA vol. 7, p. 323.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. barbata var. treleasei, D. paysonii var. treleasei
Name authority Rollins: Contr. Gray Herb. 214: 6. (1984) Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 113. (1930)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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