FNA: Draba ramulosa vs. Draba cyclomorpha

	Draba ramulosa	Draba cyclomorpha
	Tushar Mountain Draba	granite Draba, Lemmon's Draba
Habit	Perennials; (loosely matted, grayish); caudex branched (with persistent leaf bases, branches creeping, sometimes terminating in sterile rosettes); not scapose.	Perennials; (cespitose, not pulvinate); caudex branched (with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.
Stems	unbranched, 0.4–0.6 dm, densely pubescent throughout, trichomes dendritic, 3–6-rayed, (often crisped), 0.1–0.4 mm.	unbranched, 0.1–0.6(–1) dm, (thinly) pilose throughout, trichomes simple and stalked, 2- (or 3-)rayed, (often crisped), 0.1–0.8 mm.
Basal leaves	(imbricate); not rosulate; sessile; blade obovate to oblanceolate, 0.4–1.1 cm × 2–3.2 mm, margins entire, (base and margins not ciliate), surfaces pubescent with stalked, 4–8-rayed trichomes, 0.1–0.5 mm, (sometimes 1 or more rays spurred), adaxially sometimes trichomes simple.	rosulate; shortly petiolate; petiole base and margin ciliate, (trichomes simple, 0.2–1 mm); blade (somewhat fleshy), oblanceolate to obovate, 0.4–1 cm × 2–5 mm, margins entire, surfaces glabrous or hirsute, abaxially usually with stalked, 2-rayed trichomes, 0.1–0.6 mm, rarely with fewer, simple ones, (midvein obscure), adaxially with almost exclusively simple trichomes, to 0.6 mm.
Cauline leaves	2 or 3 (sometimes basal leaves spaced, flowering stem appearing to 8-leaved); sessile; blade oblong to ovate, margins entire, surfaces pubescent as basal.	0.
Racemes	4–15-flowered, ebracteate or proximalmost 1 or 2 flowers bracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem.	5–14(–20)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem.
Flowers	sepals broadly ovate, 1.7–2.4 mm, pubescent, (trichomes short-stalked, 2–5-rayed); petals yellow, obovate to oblanceolate, 3–4.5 × 1.5–2.5 mm; anthers ovate, 0.4–0.5 mm.	sepals ovate, 1.5–2.2 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, spatulate, 4–5.5 × 1.5–2.5 mm; anthers oblong, 0.6–0.7 mm.
Fruiting pedicels	divaricate-ascending, usually straight, rarely curved upward, 3–6(–10) mm, pubescent, trichomes 3–6-rayed, (crisped, 0.1–0.4 mm), and, sometimes, simple.	horizontal to divaricate-ascending, often curved upward or straight, 3–8(–10) mm, pilose as stem.
Fruits	(not appressed to rachis), ovate to elliptic, plane, flattened, 4–6.5 × 2.5–4 mm; valves densely pubescent, trichomes simple and short-stalked, 2–5-rayed, 0.08–0.35 mm; ovules 6–12 per ovary; style (0.1–)0.3–0.7 mm.	oblong to oblong-ovate, plane, flattened, 3.5–9 × 2.5–4 mm; valves glabrous; ovules 8–12 per ovary; style 0.3–1 mm.
Seeds	oblong, 1.4–1.8 × 0.8–1.2 mm.	ovoid, 1.4–1.9 × 1–1.2 mm.

	Draba ramulosa	Draba cyclomorpha
Phenology	Flowering Jun–Aug.	Flowering Jul–Aug.
Habitat	Rock outcrops, talus, gravelly soils	Basaltic talus, stony ridges
Elevation	3300-3600 m (10800-11800 ft)	2500-3100 m (8200-10200 ft)
Distribution	UT [BONAP county map]	OR [BONAP county map]
Discussion	Of conservation concern. Molecular and chromosomal data (M. D. Windham, unpubl.) strongly suggest that Draba ramulosa is an allopolyploid species. It is thought to have originated through hybridization between D. sobolifera and a member of the white-flowered, euploid lineage of M. A. Beilstein and M. D. Windham (2003). It is easily distinguished from D. sobolifera by having pale yellow to whitish (versus bright yellow) petals, grayish (versus green) foliage, non-ciliate (versus ciliate) basal leaves pubescent with 4–8-rayed (versus 2–4-rayed) trichomes, often proximally bracteate (versus ebracteate) racemes, and flattened (versus inflated basally) fruits. Draba ramulosa is known from the Tushar Mountains in south-central Utah (Beaver and Piute counties). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Draba cyclomorpha was treated by Schulz, C. L. Hitchcock (1941), and R. C. Rollins (1993) as a variety of D. lemmonii. The two taxa are quite distinct morphologically and separated by nearly 1600 kilometers, justifying their treatment as distinct species (I. A. Al-Shehbaz and M. D. Windham 2007). Draba cyclomorpha is known to us only from the Wallowa Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 7, p. 332.	FNA vol. 7, p. 305.
Parent taxa	Brassicaceae > tribe Arabideae > Draba	Brassicaceae > tribe Arabideae > Draba
Sibling taxa	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms		D. lemmonii var. cyclomorpha
Name authority	Rollins: Contr. Gray Herb. 214: 6. (1984)	Payson: Amer. J. Bot. 4: 263. (1917)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Draba cyclomorpha

Tushar Mountain Draba

granite Draba, Lemmon's Draba

Habit

Perennials; (loosely matted, grayish); caudex branched (with persistent leaf bases, branches creeping, sometimes terminating in sterile rosettes); not scapose.

Perennials; (cespitose, not pulvinate); caudex branched (with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.

Stems

unbranched, 0.4–0.6 dm, densely pubescent throughout, trichomes dendritic, 3–6-rayed, (often crisped), 0.1–0.4 mm.

unbranched, 0.1–0.6(–1) dm, (thinly) pilose throughout, trichomes simple and stalked, 2- (or 3-)rayed, (often crisped), 0.1–0.8 mm.

Basal leaves

(imbricate);

not rosulate;

sessile;

blade obovate to oblanceolate, 0.4–1.1 cm × 2–3.2 mm, margins entire, (base and margins not ciliate), surfaces pubescent with stalked, 4–8-rayed trichomes, 0.1–0.5 mm, (sometimes 1 or more rays spurred), adaxially sometimes trichomes simple.

rosulate; shortly petiolate;

petiole base and margin ciliate, (trichomes simple, 0.2–1 mm);

blade (somewhat fleshy), oblanceolate to obovate, 0.4–1 cm × 2–5 mm, margins entire, surfaces glabrous or hirsute, abaxially usually with stalked, 2-rayed trichomes, 0.1–0.6 mm, rarely with fewer, simple ones, (midvein obscure), adaxially with almost exclusively simple trichomes, to 0.6 mm.

Cauline leaves

2 or 3 (sometimes basal leaves spaced, flowering stem appearing to 8-leaved);

sessile;

blade oblong to ovate, margins entire, surfaces pubescent as basal.

Racemes

4–15-flowered, ebracteate or proximalmost 1 or 2 flowers bracteate, slightly elongated in fruit;

rachis not flexuous, pubescent as stem.

5–14(–20)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals broadly ovate, 1.7–2.4 mm, pubescent, (trichomes short-stalked, 2–5-rayed);

petals yellow, obovate to oblanceolate, 3–4.5 × 1.5–2.5 mm;

anthers ovate, 0.4–0.5 mm.

sepals ovate, 1.5–2.2 mm, pubescent, (trichomes simple and short-stalked, 2-rayed);

petals yellow, spatulate, 4–5.5 × 1.5–2.5 mm;

anthers oblong, 0.6–0.7 mm.

Fruiting pedicels

divaricate-ascending, usually straight, rarely curved upward, 3–6(–10) mm, pubescent, trichomes 3–6-rayed, (crisped, 0.1–0.4 mm), and, sometimes, simple.

horizontal to divaricate-ascending, often curved upward or straight, 3–8(–10) mm, pilose as stem.

Fruits

(not appressed to rachis), ovate to elliptic, plane, flattened, 4–6.5 × 2.5–4 mm;

valves densely pubescent, trichomes simple and short-stalked, 2–5-rayed, 0.08–0.35 mm;

ovules 6–12 per ovary;

style (0.1–)0.3–0.7 mm.

oblong to oblong-ovate, plane, flattened, 3.5–9 × 2.5–4 mm;

valves glabrous;

ovules 8–12 per ovary;

style 0.3–1 mm.

Seeds

oblong, 1.4–1.8 × 0.8–1.2 mm.

ovoid, 1.4–1.9 × 1–1.2 mm.

Draba ramulosa

Draba cyclomorpha

Phenology

Flowering Jun–Aug.

Flowering Jul–Aug.

Habitat

Rock outcrops, talus, gravelly soils

Basaltic talus, stony ridges

Elevation

3300-3600 m (10800-11800 ft)

2500-3100 m (8200-10200 ft)

Distribution

[BONAP county map]

Discussion

Of conservation concern.

Molecular and chromosomal data (M. D. Windham, unpubl.) strongly suggest that Draba ramulosa is an allopolyploid species. It is thought to have originated through hybridization between D. sobolifera and a member of the white-flowered, euploid lineage of M. A. Beilstein and M. D. Windham (2003). It is easily distinguished from D. sobolifera by having pale yellow to whitish (versus bright yellow) petals, grayish (versus green) foliage, non-ciliate (versus ciliate) basal leaves pubescent with 4–8-rayed (versus 2–4-rayed) trichomes, often proximally bracteate (versus ebracteate) racemes, and flattened (versus inflated basally) fruits. Draba ramulosa is known from the Tushar Mountains in south-central Utah (Beaver and Piute counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba cyclomorpha was treated by Schulz, C. L. Hitchcock (1941), and R. C. Rollins (1993) as a variety of D. lemmonii. The two taxa are quite distinct morphologically and separated by nearly 1600 kilometers, justifying their treatment as distinct species (I. A. Al-Shehbaz and M. D. Windham 2007). Draba cyclomorpha is known to us only from the Wallowa Mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 7, p. 332.

FNA vol. 7, p. 305.

Parent taxa

Brassicaceae > tribe Arabideae > Draba

Sibling taxa

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

Synonyms

D. lemmonii var. cyclomorpha

Name authority

Rollins: Contr. Gray Herb. 214: 6. (1984)

Payson: Amer. J. Bot. 4: 263. (1917)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Flora of North America species comparison

Draba ramulosa

Draba cyclomorpha

Draba ramulosa

Draba cyclomorpha