Draba praealta |
Draba sobolifera |
|
---|---|---|
Draba, tall Draba, tall whitlow-grass, tall whitlow-grass Draba |
stolon Draba |
|
Habit | Annuals or perennials; (short-lived); caudex simple or branched; not scapose. | Perennials; (cespitose); caudex often branched (with persistent leaf bases, branches elongated, sometimes terminating in sterile rosettes); scapose. |
Stems | branched or unbranched, (0.5–)0.8–3.2(–3.8) dm, pubescent throughout, trichomes 3–5-rayed, 0.1–0.4 mm, with simple and 2-rayed ones, 0.4–1 mm. |
unbranched, (0.15–)0.25–0.6(–0.7) dm, pubescent basally, trichomes simple and 2- or 3-rayed, 0.1–0.5 mm. |
Basal leaves | rosulate; subsessile; petiole base ciliate, (trichomes simple and 2-rayed, 0.2–0.8 mm); blade oblanceolate, (0.7–)1–3.5(–4.5) cm × 1.5–7(–9) mm, margins usually dentate, rarely entire, surfaces pubescent, abaxially with stalked, 3–6-rayed trichomes, 0.1–0.5 mm, adaxially with stalked, 3–5-rayed, 0.1–0.6 mm, and simple ones, to 0.8 mm. |
rosulate; subsessile; petiole (0–0.5 cm) margin ciliate, (trichomes simple, 0.2–1 mm); blade oblanceolate to spatulate or obovate, (0.3–)0.4–1.4(–2) cm × (1.5–)2–4(–6) mm, margins entire, (ciliate as petiole), surfaces usually pubescent, sometimes glabrescent, abaxially with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, adaxially often with simple and 2-rayed trichomes, to 0.9 mm. |
Cauline leaves | (1 or) 2–5(–9); sessile; blade ovate to lanceolate, margins dentate or entire, surfaces pubescent as basal. |
0 (or 1); sessile; blade similar to basal. |
Racemes | (5–)8–30(–37)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
(5–)9–18(–23)-flowered, ebracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stems proximally (trichomes often crisped). |
Flowers | (late-season ones petaliferous); sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals white, oblanceolate to spatulate, 2.8–3.5(–4) × 0.8–1.2 mm; anthers ovate, 0.2–0.3 mm. |
sepals ovate or broadly oblong, 1.7–2.5 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, spatulate to oblanceolate, 3–4 × 1.5–2 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | divaricate-ascending to ascending or suberect (not appressed to rachis), straight, (3–)4–10(–12) mm, pubescent, trichomes 3–5-rayed (0.1–0.4 mm), and sometimes simple. |
divaricate-ascending, usually straight, rarely curved upward, 3–6(–10) mm, pubescent, trichomes simple and 2- or 3-rayed, (crisped, 0.1–0.5 mm). |
Fruits | (erect, not appressed to rachis) lanceolate to linear-lanceolate, plane, flattened, (5–)7–12(–15) × 1.5–2.5(–3) mm; valves puberulent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.25 mm; ovules (24–)30–52 per ovary; style 0.03–0.1(0.15) mm. |
ovoid or ovoid-lanceolate, plane, inflated at least basally, (3–)4–6(–7) × 2–4 mm; valves glabrous or pubescent, trichomes simple and 2-rayed, 0.07–0.3 mm; ovules (4–)8–12 per ovary; style (0.3–)0.4–0.6 mm. |
Seeds | oblong, 0.8–1.1 × 0.5–0.6 mm, (often apiculate). |
oblong, 1–1.4 × 0.6–0.8 mm. |
2n | = 56. |
= 26. |
Draba praealta |
Draba sobolifera |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Rocky pine slopes, subalpine areas, alpine meadows, grassy volcanic ridges, woodlands, damp rocky areas, limestone talus, shale cliffs, steep hillsides | Igneous rock outcrops, fellfields, rocky slopes in mixed conifer and alpine meadow communities |
Elevation | 1200-2900 m (3900-9500 ft) | 3100-3600 m (10200-11800 ft) |
Distribution |
CA; ID; MT; NV; OR; WA; WY; AB; BC; NT; YT; e Asia (Russian Far East [Chukchi Peninsula])
|
UT |
Discussion | Morphological and chromosomal evidence (M. D. Windham, unpubl.) suggests that Draba praealta may be an allopolyploid originating through hybridization between D. albertina and D. cana. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) placed D. lonchocarpa var. dasycarpa in the synonymy of D. lonchocarpa. G. A. Mulligan (1974) indicated that the type of var. dasycarpa does not belong to that species and he correctly annotated its isotype (Macoun 64454, CAN) as D. praealta. Mulligan (1976) and Rollins indicated that D. praealta occurs in Alaska, but we have not seen unequivocal material from the state. We have not seen the material on which A. E. Katenin and V. V. Petrovsky (1995) based their record from the Russian Far East. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba sobolifera is related to, and sympatric with, D. ramulosa, and occasional sterile hybrids are encountered. There is no evidence of introgression and the two species are easily distinguished (see discussion of 90. D. ramulosa). Draba sobolifera is known from the Tushar Mountains in south-central Utah (Beaver and Piute counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 331. | FNA vol. 7, p. 338. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. cascadensis, D. columbiana, D. dolichopoda, D. lapilutea, D. lonchocarpa var. dasycarpa, D. praealta var. yellowstonensis, D. yellowstonensis | D. sobolifera var. uncinalis, D. uncinalis |
Name authority | Greene: Pittonia 3: 306. (1898) | Rydberg: Bull. Torrey Bot. Club 30: 251. (1903) |
Web links |
|