Draba praealta |
Draba pennellii |
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Draba, tall Draba, tall whitlow-grass, tall whitlow-grass Draba |
schell creek Draba |
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Habit | Annuals or perennials; (short-lived); caudex simple or branched; not scapose. | Perennials; (cespitose, often forming mats); caudex branched (elongated, with persistent leaf bases, branches sometimes terminating in sterile rosettes); sometimes scapose. |
Stems | branched or unbranched, (0.5–)0.8–3.2(–3.8) dm, pubescent throughout, trichomes 3–5-rayed, 0.1–0.4 mm, with simple and 2-rayed ones, 0.4–1 mm. |
unbranched, (0.15–)0.3–0.7(–1) dm, pubescent proximally, trichomes simple and spurred, 0.4–0.8 mm, and short-stalked, 2–4-rayed, 0.1–0.4 mm, (sometimes with simple trichomes distally). |
Basal leaves | rosulate; subsessile; petiole base ciliate, (trichomes simple and 2-rayed, 0.2–0.8 mm); blade oblanceolate, (0.7–)1–3.5(–4.5) cm × 1.5–7(–9) mm, margins usually dentate, rarely entire, surfaces pubescent, abaxially with stalked, 3–6-rayed trichomes, 0.1–0.5 mm, adaxially with stalked, 3–5-rayed, 0.1–0.6 mm, and simple ones, to 0.8 mm. |
subrosulate; sessile; blade oblanceolate to oblong, (0.3–)0.5–0.8 cm × 1–3 mm, (base ciliate, trichomes simple, 0.2–0.8 mm), margins entire, (not ciliate), surfaces usually pubescent, with stalked, (2–)4 (or 5)-rayed trichomes, 0.2–0.5 mm, adaxially sometimes glabrate, or primarily with simple trichomes, 0.3–0.8 mm. |
Cauline leaves | (1 or) 2–5(–9); sessile; blade ovate to lanceolate, margins dentate or entire, surfaces pubescent as basal. |
(0 or) 1–4; sessile; blade ovate to oblong, margins entire, surfaces pubescent as basal. |
Racemes | (5–)8–30(–37)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
(3–)7–15(–20)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | (late-season ones petaliferous); sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals white, oblanceolate to spatulate, 2.8–3.5(–4) × 0.8–1.2 mm; anthers ovate, 0.2–0.3 mm. |
sepals ovate to oblong, 2–3 mm, pubescent, (trichomes simple, with fewer, short-stalked, 2-rayed ones); petals white, obovate, 3.5–6 × 1.5–2.8 mm; anthers oblong, 0.5–0.6 mm. |
Fruiting pedicels | divaricate-ascending to ascending or suberect (not appressed to rachis), straight, (3–)4–10(–12) mm, pubescent, trichomes 3–5-rayed (0.1–0.4 mm), and sometimes simple. |
divaricate-ascending to ascending, straight, 3–10 mm, pubescent, trichomes simple and short-stalked, 2–4-rayed, (0.1–0.4 mm). |
Fruits | (erect, not appressed to rachis) lanceolate to linear-lanceolate, plane, flattened, (5–)7–12(–15) × 1.5–2.5(–3) mm; valves puberulent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.25 mm; ovules (24–)30–52 per ovary; style 0.03–0.1(0.15) mm. |
broadly or, rarely, narrowly lanceolate to elliptic or ovate, plane or slightly twisted, flattened, (4)5–8(–10) × 2–3.2 mm; valves (each with distinct midvein), pubescent, trichomes simple and short-stalked, 2-rayed, 0.08–0.2 mm; ovules 12–24 per ovary; style (0.7–)1–1.8(–2.1) mm. |
Seeds | oblong, 0.8–1.1 × 0.5–0.6 mm, (often apiculate). |
oblong, 0.9–1.3 × 0.7–1 mm. |
2n | = 56. |
= 32. |
Draba praealta |
Draba pennellii |
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Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Rocky pine slopes, subalpine areas, alpine meadows, grassy volcanic ridges, woodlands, damp rocky areas, limestone talus, shale cliffs, steep hillsides | Rock outcrops and talus slopes in pinyon-juniper, sagebrush, mountain shrub, and mixed conifer communities |
Elevation | 1200-2900 m (3900-9500 ft) | 1900-3500 m (6200-11500 ft) |
Distribution |
CA; ID; MT; NV; OR; WA; WY; AB; BC; NT; YT; e Asia (Russian Far East [Chukchi Peninsula])
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NV |
Discussion | Morphological and chromosomal evidence (M. D. Windham, unpubl.) suggests that Draba praealta may be an allopolyploid originating through hybridization between D. albertina and D. cana. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) placed D. lonchocarpa var. dasycarpa in the synonymy of D. lonchocarpa. G. A. Mulligan (1974) indicated that the type of var. dasycarpa does not belong to that species and he correctly annotated its isotype (Macoun 64454, CAN) as D. praealta. Mulligan (1976) and Rollins indicated that D. praealta occurs in Alaska, but we have not seen unequivocal material from the state. We have not seen the material on which A. E. Katenin and V. V. Petrovsky (1995) based their record from the Russian Far East. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba pennellii is known from White Pine County in east-central Nevada. It is easily distinguished from other white-flowered species in the region by its elongated, many-branched caudices, long styles, usually few-leaved or, rarely, leafless flowering stems, and distinct midvein on fruit valves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 331. | FNA vol. 7, p. 329. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. cascadensis, D. columbiana, D. dolichopoda, D. lapilutea, D. lonchocarpa var. dasycarpa, D. praealta var. yellowstonensis, D. yellowstonensis | |
Name authority | Greene: Pittonia 3: 306. (1898) | Rollins: J. Arnold Arbor. 64: 502. (1983) |
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