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Draba, tall Draba, tall whitlow-grass, tall whitlow-grass Draba

schell creek Draba

Habit Annuals or perennials; (short-lived); caudex simple or branched; not scapose. Perennials; (cespitose, often forming mats); caudex branched (elongated, with persistent leaf bases, branches sometimes terminating in sterile rosettes); sometimes scapose.
Stems

branched or unbranched, (0.5–)0.8–3.2(–3.8) dm, pubescent throughout, trichomes 3–5-rayed, 0.1–0.4 mm, with simple and 2-rayed ones, 0.4–1 mm.

unbranched, (0.15–)0.3–0.7(–1) dm, pubescent proximally, trichomes simple and spurred, 0.4–0.8 mm, and short-stalked, 2–4-rayed, 0.1–0.4 mm, (sometimes with simple trichomes distally).

Basal leaves

rosulate;

subsessile;

petiole base ciliate, (trichomes simple and 2-rayed, 0.2–0.8 mm);

blade oblanceolate, (0.7–)1–3.5(–4.5) cm × 1.5–7(–9) mm, margins usually dentate, rarely entire, surfaces pubescent, abaxially with stalked, 3–6-rayed trichomes, 0.1–0.5 mm, adaxially with stalked, 3–5-rayed, 0.1–0.6 mm, and simple ones, to 0.8 mm.

subrosulate;

sessile;

blade oblanceolate to oblong, (0.3–)0.5–0.8 cm × 1–3 mm, (base ciliate, trichomes simple, 0.2–0.8 mm), margins entire, (not ciliate), surfaces usually pubescent, with stalked, (2–)4 (or 5)-rayed trichomes, 0.2–0.5 mm, adaxially sometimes glabrate, or primarily with simple trichomes, 0.3–0.8 mm.

Cauline leaves

(1 or) 2–5(–9);

sessile;

blade ovate to lanceolate, margins dentate or entire, surfaces pubescent as basal.

(0 or) 1–4;

sessile;

blade ovate to oblong, margins entire, surfaces pubescent as basal.

Racemes

(5–)8–30(–37)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

(3–)7–15(–20)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

(late-season ones petaliferous);

sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and stalked, 2-rayed);

petals white, oblanceolate to spatulate, 2.8–3.5(–4) × 0.8–1.2 mm;

anthers ovate, 0.2–0.3 mm.

sepals ovate to oblong, 2–3 mm, pubescent, (trichomes simple, with fewer, short-stalked, 2-rayed ones);

petals white, obovate, 3.5–6 × 1.5–2.8 mm;

anthers oblong, 0.5–0.6 mm.

Fruiting pedicels

divaricate-ascending to ascending or suberect (not appressed to rachis), straight, (3–)4–10(–12) mm, pubescent, trichomes 3–5-rayed (0.1–0.4 mm), and sometimes simple.

divaricate-ascending to ascending, straight, 3–10 mm, pubescent, trichomes simple and short-stalked, 2–4-rayed, (0.1–0.4 mm).

Fruits

(erect, not appressed to rachis) lanceolate to linear-lanceolate, plane, flattened, (5–)7–12(–15) × 1.5–2.5(–3) mm;

valves puberulent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.25 mm;

ovules (24–)30–52 per ovary;

style 0.03–0.1(0.15) mm.

broadly or, rarely, narrowly lanceolate to elliptic or ovate, plane or slightly twisted, flattened, (4)5–8(–10) × 2–3.2 mm;

valves (each with distinct midvein), pubescent, trichomes simple and short-stalked, 2-rayed, 0.08–0.2 mm;

ovules 12–24 per ovary;

style (0.7–)1–1.8(–2.1) mm.

Seeds

oblong, 0.8–1.1 × 0.5–0.6 mm, (often apiculate).

oblong, 0.9–1.3 × 0.7–1 mm.

2n

= 56.

= 32.

Draba praealta

Draba pennellii

Phenology Flowering Jun–Aug. Flowering May–Jul.
Habitat Rocky pine slopes, subalpine areas, alpine meadows, grassy volcanic ridges, woodlands, damp rocky areas, limestone talus, shale cliffs, steep hillsides Rock outcrops and talus slopes in pinyon-juniper, sagebrush, mountain shrub, and mixed conifer communities
Elevation 1200-2900 m (3900-9500 ft) 1900-3500 m (6200-11500 ft)
Distribution
from FNA
CA; ID; MT; NV; OR; WA; WY; AB; BC; NT; YT; e Asia (Russian Far East [Chukchi Peninsula])
[WildflowerSearch map]
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Morphological and chromosomal evidence (M. D. Windham, unpubl.) suggests that Draba praealta may be an allopolyploid originating through hybridization between D. albertina and D. cana. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) placed D. lonchocarpa var. dasycarpa in the synonymy of D. lonchocarpa. G. A. Mulligan (1974) indicated that the type of var. dasycarpa does not belong to that species and he correctly annotated its isotype (Macoun 64454, CAN) as D. praealta. Mulligan (1976) and Rollins indicated that D. praealta occurs in Alaska, but we have not seen unequivocal material from the state.

We have not seen the material on which A. E. Katenin and V. V. Petrovsky (1995) based their record from the Russian Far East.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Draba pennellii is known from White Pine County in east-central Nevada. It is easily distinguished from other white-flowered species in the region by its elongated, many-branched caudices, long styles, usually few-leaved or, rarely, leafless flowering stems, and distinct midvein on fruit valves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 331. FNA vol. 7, p. 329.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. cascadensis, D. columbiana, D. dolichopoda, D. lapilutea, D. lonchocarpa var. dasycarpa, D. praealta var. yellowstonensis, D. yellowstonensis
Name authority Greene: Pittonia 3: 306. (1898) Rollins: J. Arnold Arbor. 64: 502. (1983)
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