Draba petrophila |
Draba cinerea |
|
---|---|---|
Santa Rita Mountain Draba, Santa Rita Mountain whitlowgrass, Santa Rita whitlow grass |
gray-leaf Draba, gray-leaf whitlow-grass |
|
Habit | Perennials; (cespitose, long-lived); caudex simple or branched (covered with persistent petioles), not scapose. | Perennials; (cespitose); caudex simple or branched; rarely scapose. |
Stems | usually unbranched, rarely branched, 0.3–1.9(–2.8) dm, moderately to densely hirsute throughout, trichomes simple, 0.2–1.3 mm, with short-stalked to subsessile, cruciform, 0.02–0.4 mm, and 2-rayed ones, 0.3–0.9 mm. |
unbranched, (0.3–)0.5–1.6(–2.7) dm, pubescent throughout, trichomes 4–10-rayed, 0.1–0.3 mm, (often some rays branched, simple trichomes sparse, to 0.6 mm). |
Basal leaves | rosulate; petiolate; petiole ciliate, (trichomes simple, 0.4–1.5 mm); blade oblanceolate, 1–5(–6) cm × 2–5(–10) mm, margins usually entire, rarely denticulate, surfaces pubescent, abaxially with stalked, cruciform trichomes, 0.07–0.5 mm, adaxially often similar, sometimes with fewer, simple and 2-rayed trichomes, 0.4–1.3 mm. |
rosulate; petiolate; petiole base and proximalmost margin ciliate, (trichomes simple, 0.2–0.8 mm); blade oblanceolate to narrowly obovate or linear-lanceolate, 0.4–1.5 cm × 1–5 mm, margins usually entire, rarely with 1 tooth on each side, surfaces densely pubescent with minutely stalked, stellate, 8–12-rayed, trichomes 0.1–0.25 mm, (midvein obscure abaxially, apex trichomes simple). |
Cauline leaves | 3–10; sessile; blade ovate to lanceolate or oblong, margins entire or denticulate, surfaces pubescent as basal. |
0–3(–5); sessile; blade ovate or oblong to lanceolate, margins entire, surfaces pubescent as basal. |
Racemes | 10–37(–58)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
(3–)5–17(–24)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 2–3.5 mm, pubescent, (trichomes simple and 2-rayed); petals yellow, oblanceolate, 3.5–6 × 1.2–1.8 mm; anthers oblong, 0.8–1 mm. |
sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and short-stalked, 2- or 3-rayed); petals white, spatulate to obovate, 3.5–4.5 × 1.5–2 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | divaricate-ascending, straight, 4–10 mm, pubescent throughout, trichomes simple (0.1–0.5 mm) and subsessile, 2–4-rayed, (0.03–0.2 mm). |
divaricate-ascending or ascending, straight, (3–)4–7(–9) mm, pubescent as stem. |
Fruits | lanceolate to elliptic, often strongly twisted, flattened, 5–11 × 2–3 mm; valves puberulent at least along margin, trichomes simple, antrorse, 0.03–0.15 mm; ovules 14–24 per ovary; style 0.8–1.8(–2.5) mm. |
oblong to elliptic, plane, slightly flattened, 5–8 × 2–3 mm; valves pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.3 mm, (some rays branched); ovules 20–36(–42) per ovary; style (0.1–)0.2–1 mm. |
Seeds | ovoid, 1–1.4 × 0.6–0.9 mm. |
ovoid, 0.6–0.8 × 0.4–0.6 mm. |
2n | = 48. |
|
Draba petrophila |
Draba cinerea |
|
Phenology | Flowering Jul–Sep. | Flowering Jun–Jul. |
Habitat | Crevices, ledges of cliffs | Rock outcrops, ridges, meadows, gravel beaches, stream banks, alluvial fans |
Elevation | 1200-2800 m (3900-9200 ft) | 0-500 m (0-1600 ft) |
Distribution |
AZ
|
AK; BC; NT; NU; ON; QC; SK; YT; Greenland; Europe (Finland, Norway, n Russia); e Asia (Russian Far East, Siberia) |
Discussion | Draba petrophila is often broadly circumscribed to include the taxon herein called D. viridis. For a discussion of the differences between these species and the closely related D. helleriana, see I. A. Al-Shehbaz and M. D. Windham (2007). Draba petrophila, in the strict sense, is known only from Cochise, Pima, and Santa Cruz counties in southeastern Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba cinerea is a polymorphic species in which tetraploid, hexaploid, heptaploid, and octoploid chromosome counts (all based on x = 8) have been reported (e.g., R. C. Rollins 1993). Although we have not seen the type collection, we tentatively consider the taxon to be hexaploid, which is the predominant ploidy level throughout the species range, including North America, and also from its type region in Siberia. Hexaploid Draba cinerea is closely related to decaploid (2n = 80) D. arctica, from which it is distinguished with difficulty. In general, D. cinerea has 1–3(–5)-leaved stems, basal leaf blades usually not terminated with simple trichomes and with obscure midveins abaxially, petals 3.5–4.5 mm, and seeds 0.6–0.8 × 0.4–0.6 mm. By contrast, D. arctica has leafless or 1(–3)-leaved stems, basal leaf blades terminated with simple or 2-rayed trichomes and with midveins distinct abaxially, petals 3.5–6 mm, and seeds (0.8–)0.9–1.1 × (0.6–)0.7–0.8 mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 329. | FNA vol. 7, p. 300. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. helleriana var. blumeri, D. helleriana var. petrophila | D. magellanica subsp. cinerea |
Name authority | Greene: Pittonia 4: 17. (1899) | Adams: Mém Soc. Imp. Naturalistes Moscou 5: 103. (1817) |
Web links |