Draba pectinipila |
Draba zionensis |
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Habit | Perennials; (cespitose, sometimes forming mats); caudex branched (with persistent leaf bases, branches creeping, terminating in scapes or sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.3–)0.4–1.6(–1.9) dm, pubescent, trichomes sessile, pectinate, 0.1–0.4 mm, (parallel to long axis of stem, sometimes with irregularly 2–4-rayed ones, 0.2–0.6 mm). |
unbranched, 0.5–1.5(–1.8) dm, usually sparsely pubescent proximally, usually glabrous distally, rarely throughout, trichomes simple, 0.3–0.8 mm, often with smaller, 2–4-rayed ones. |
Basal leaves | rosulate; subsessile; blade narrowly oblanceolate to linear, (0.4–)0.6–1.3(–2) cm × 0.9–2.2 mm, margins entire (not ciliate), surfaces pubescent with subsessile or sessile, pectinate trichomes, 0.2–0.5 mm. |
rosulate; shortly petiolate; petiole base sometimes ciliate (margin not ciliate, trichomes simple and 2-rayed, 0.2–0.5 mm); blade spatulate to obovate, 0.7–3.5 cm × 3–10(–12) mm, margins usually entire, rarely obscurely dentate (near apex), surfaces sparsely pubescent with stalked, (2–)4 (or 5)-rayed trichomes, 0.05–0.5 mm, (midvein obscure). |
Cauline leaves | 0. |
0. |
Racemes | 5–22-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
14–36-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals broadly ovate, 2–3.2 mm, pubescent, (trichomes pectinate); petals yellow, obovate to spatulate, 4–6.5 × 1.5–2.5 mm; anthers ovate 0.4–0.5 mm. |
sepals ovate, 2–3 mm, glabrous or sparsely pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals orange-yellow (fading pale yellow), broadly spatulate to subovate, 5–6 × 1.8–2.8 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | divaricate-ascending, straight, (5–)7–14 mm, sparsely pubescent, trichomes pectinate. |
ascending to divaricate-ascending, straight, (not expanded basally), (4–)5–12(–15) mm, glabrous. |
Fruits | ovoid, plane, slightly inflated basally, 4–6(–7) × 2–3 mm; valves pubescent, trichomes usually sessile, pectinate, 0.2–0.5 mm, rarely with simple ones; ovules 4–8 per ovary; style 0.5–1.5 mm. |
lanceolate to linear-lanceolate or ovate, plane (not curved), flattened, 6–12(–17) × 2–3 mm; valves glabrous; ovules 12–20 per ovary; style 0.4–1 mm. |
Seeds | oblong, 1.1–1.5 × 0.6–0.8 mm. |
oblong, 1–1.3 × 0.6–0.8 mm. |
2n | = 22. |
= 26. |
Draba pectinipila |
Draba zionensis |
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Phenology | Flowering Apr–Jun. | Flowering Mar–May. |
Habitat | Rocky slopes in sagebrush scrub and pinyon-juniper woodlands | Sandstone (rarely limestone) rock outcrops and sandy slopes in pinyon-juniper or pine communities |
Elevation | 1700-2400 m (5600-7900 ft) | 1000-2500 m (3300-8200 ft) |
Distribution |
CO; MT; UT; WY |
UT |
Discussion | Draba pectinipila was treated as a synonym of D. oligosperma by G. A. Mulligan (1972), R. C. Rollins (1993), and N. H. Holmgren (2005b). The latter recognized D. juniperina as a distinct species but overlooked the fact that it is indistinguishable from the type collections of the earlier-published D. pectinipila. The species (including D. juniperina) differs significantly from D. oligosperma in both chromosome number and morphology. Draba pectinipila is easily distinguished by having fruit valves pubescent with pectinate trichomes, fruiting pedicels (5–)7–14 mm, petals 4–6.5 mm, ovules 4–8 per ovary, and styles 0.5–1.5 mm. By contrast, D. oligosperma has fruit valves glabrous or pubescent with simple or 2-rayed trichomes, fruiting pedicels (2–)3–10(–13) mm, petals 2.5–4 mm, ovules 6–12 per ovary, and styles 0.1–0.8(–1.1) mm. Draba pectinipila was previously known only from the type locality in northwestern Wyoming (Park County). Its range is now expanded to include that of D. juniperina in northwestern Colorado (Moffat County), northeastern Utah (Daggett and Uintah counties), and southwestern Wyoming (Sweetwater County). The record from Uintah County is based on Goodrich 22275 (NY). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
R. C. Rollins (1993) treated Draba zionensis as a variety of D. asprella, but its true relationships appear to lie with two other southern Utah endemics, D. sobolifera and D. subalpina. Draba zionensis is easily distinguished from D. subalpina by having orange-yellow (versus white) petals, and from D. asprella and D. sobolifera by its glabrous (versus pubescent) pedicels and stems distally. Nearly all populations of the species are found in and around Zion National Park in southwestern Utah (Iron, Kane, and Washington counties). A specimen supposedly from the Deep Creek Mountains (Juab County) may be mislabeled. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 328. | FNA vol. 7, p. 346. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. juniperina, D. oligosperma var. juniperina, D. oligosperma var. pectinipila | D. asprella var. zionensis |
Name authority | Rollins: Rhodora 55: 231. (1953) | C. L. Hitchcock: Revis. Drabas W. N. Amer., 49, plate 2, fig. 16. (1941) |
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