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bitter root Draba

Habit Perennials; (cespitose, sometimes forming mats); caudex branched (with persistent leaf bases, branches creeping, terminating in scapes or sterile rosettes); scapose. Perennials; (densely pulvinate); caudex branched (branches elongated, loose, with persistent leaf remains, terminating in flowering or sterile shoots); scapose.
Stems

unbranched, (0.3–)0.4–1.6(–1.9) dm, pubescent, trichomes sessile, pectinate, 0.1–0.4 mm, (parallel to long axis of stem, sometimes with irregularly 2–4-rayed ones, 0.2–0.6 mm).

unbranched, (0.05–)0.2–0.6 dm, glabrous.

Basal leaves

rosulate;

subsessile;

blade narrowly oblanceolate to linear, (0.4–)0.6–1.3(–2) cm × 0.9–2.2 mm, margins entire (not ciliate), surfaces pubescent with subsessile or sessile, pectinate trichomes, 0.2–0.5 mm.

(densely imbricate); rosulate;

petiolate;

petiole ciliate throughout;

blade (fleshy), oblong to obovate or oblanceolate, 0.3–0.7(–1) cm × 1–2(–2.5) mm, margins entire, (ciliate, trichomes simple, 0.1–0.5 mm, apex obtuse), surfaces glabrous (midvein obscure abaxially).

Cauline leaves

0.

0.

Racemes

5–22-flowered, ebracteate, considerably elongated in fruit;

rachis not flexuous, pubescent as stem.

2–8(–10)-flowered, ebracteate, (subcorymbose), slightly elongated in fruit;

rachis not flexuous, glabrous.

Flowers

sepals broadly ovate, 2–3.2 mm, pubescent, (trichomes pectinate);

petals yellow, obovate to spatulate, 4–6.5 × 1.5–2.5 mm;

anthers ovate 0.4–0.5 mm.

sepals oblong, 1.5–2.2 mm, glabrous;

petals pale to bright yellow, spatulate, 3.5–4 × 1–2 mm;

anthers ovate, 0.3–0.4 mm.

Fruiting pedicels

divaricate-ascending, straight, (5–)7–14 mm, sparsely pubescent, trichomes pectinate.

divaricate-ascending (not decurrent basally), straight, 4–10 mm, glabrous.

Fruits

ovoid, plane, slightly inflated basally, 4–6(–7) × 2–3 mm;

valves pubescent, trichomes usually sessile, pectinate, 0.2–0.5 mm, rarely with simple ones;

ovules 4–8 per ovary;

style 0.5–1.5 mm.

ovate to oblong-elliptic, plane, flattened, 4–8 × 2–4 mm;

valves (obscurely veined), glabrous;

ovules 6–14 per ovary;

style 0.1–0.5 mm.

Seeds

oblong, 1.1–1.5 × 0.6–0.8 mm.

ovoid, 1.2–1.5 × 0.8–1 mm.

2n

= 22.

Draba pectinipila

Draba daviesiae

Phenology Flowering Apr–Jun. Flowering Jul–Aug.
Habitat Rocky slopes in sagebrush scrub and pinyon-juniper woodlands Talus slopes, rock crevices and cracks, rocky ridges and slides, alpine meadows
Elevation 1700-2400 m (5600-7900 ft) 2700-2900 m (8900-9500 ft)
Distribution
from FNA
CO; MT; UT; WY
[BONAP county map]
from FNA
MT
[BONAP county map]
Discussion

Draba pectinipila was treated as a synonym of D. oligosperma by G. A. Mulligan (1972), R. C. Rollins (1993), and N. H. Holmgren (2005b). The latter recognized D. juniperina as a distinct species but overlooked the fact that it is indistinguishable from the type collections of the earlier-published D. pectinipila. The species (including D. juniperina) differs significantly from D. oligosperma in both chromosome number and morphology. Draba pectinipila is easily distinguished by having fruit valves pubescent with pectinate trichomes, fruiting pedicels (5–)7–14 mm, petals 4–6.5 mm, ovules 4–8 per ovary, and styles 0.5–1.5 mm. By contrast, D. oligosperma has fruit valves glabrous or pubescent with simple or 2-rayed trichomes, fruiting pedicels (2–)3–10(–13) mm, petals 2.5–4 mm, ovules 6–12 per ovary, and styles 0.1–0.8(–1.1) mm. Draba pectinipila was previously known only from the type locality in northwestern Wyoming (Park County). Its range is now expanded to include that of D. juniperina in northwestern Colorado (Moffat County), northeastern Utah (Daggett and Uintah counties), and southwestern Wyoming (Sweetwater County). The record from Uintah County is based on Goodrich 22275 (NY).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although originally described as a variety of Draba apiculata (= D. globosa), D. daviesiae is distinct morphologically. It is easily distinguished from the former by its densely pulvinate habit, obtuse leaf blades, and obscurely veined fruit valves. By contrast, D. globosa exhibits a cespitose but non-pulvinate habit, acute leaf blades, and prominently veined fruit valves. Draba daviesiae is known from the Bitterroot Mountains in Ravalli County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 328. FNA vol. 7, p. 305.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. juniperina, D. oligosperma var. juniperina, D. oligosperma var. pectinipila D. apiculata var. daviesiae, D. densifolia var. daviesiae
Name authority Rollins: Rhodora 55: 231. (1953) (C. L. Hitchcock) Rollins: Contr. Gray Herb. 214: 5. (1984)
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