Draba oreibata |
Draba spectabilis |
|
---|---|---|
limestone Draba |
showy Draba, showy whitlowgrass |
|
Habit | Perennials; (cespitose); caudex branched (with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. | Perennials; caudex simple or branched (sometimes with persistent leaf bases); not scapose. |
Stems | unbranched, 0.2–0.9 dm, glabrous throughout. |
unbranched, (0.7–)1.1–3.7(–5.3) dm, pubescent throughout (sparsely so distally), trichomes simple, malpighiaceous, or sessile and 4-rayed, 0.15–0.5 mm, (2 rays parallel to stem axis longer). |
Basal leaves | rosulate; sessile; blade oblong to narrowly oblanceolate, 0.2–1 cm × 0.6–2(–4) mm, margins entire, (ciliate, trichomes simple, 0.1–0.5 mm, apex obtuse), surfaces glabrous. |
rosulate; petiolate; petiole sometimes ciliate, (trichomes simple); blade oblanceolate to spatulate or obovate, (1–)1.5–4.4(–6.3) cm × (3–)5–12(–15) mm, margins entire or dentate, surfaces pubescent with sessile, often 4-rayed trichomes, 0.1–0.6 mm (longest rays parallel to midvein, lateral rays sometimes reduced to tiny spurs), sometimes malpighiaceous trichomes present. |
Cauline leaves | 0. |
4–12(–17); sessile; blade broadly ovate to lanceolate or oblong, margins entire or dentate, (ciliate or not), surfaces pubescent as basal or adaxially also with simple trichomes. |
Racemes | 3–8-flowered, ebracteate, considerably elongated in fruit; rachis slightly flexuous, glabrous. |
(10–)16–49(–61)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 1.7–2.5 mm, glabrous; petals white, oblanceolate to obovate, 3.5–4.5 × 1.2–2 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, (2.2–)2.5–4 mm, glabrous or pubescent, (trichomes simple and malpighiaceous); petals yellow, oblanceolate, 4–6.5 × 1–2.5 mm; anthers oblong, 0.5–0.7 mm. |
Fruiting pedicels | divaricate-ascending to ascending, straight, 4–13 mm, glabrous. |
horizontal to divaricate-ascending, straight or curved upward, (5–)7–20(–26) mm, glabrous or pubescent as rachis abaxially. |
Fruits | oblong to narrowly so, slightly twisted, flattened, 5–9 × 1.5–3 mm; valves glabrous; ovules 8–16 per ovary; style 0.3–0.9(–1.2) mm. |
lanceolate to oblong or elliptic, plane, flattened, (6–)7–13 × 2–3.5 mm; valves glabrous or puberulent, trichomes simple and sessile, 2-rayed, 0.03–0.3 mm; ovules 12–24 per ovary; style (0.5–)1–2.7 mm. |
Seeds | oblong, 0.9–1.3 × 0.6–0.8 mm. |
ovoid, 1–1.4 × 0.7–0.9 mm. |
2n | = 32. |
= 40. |
Draba oreibata |
Draba spectabilis |
|
Phenology | Flowering May–Jul. | Flowering Jun–Aug. |
Habitat | Limestone cliffs, talus | Talus, rocky hillsides, meadows in open conifer forests, aspen groves, and alpine communities |
Elevation | 1800-2700 m (5900-8900 ft) | 2000-3900 m (6600-12800 ft) |
Distribution |
ID |
CO; NM; WY
|
Discussion | R. C. Rollins (1993) and N. H. Holmgren (2005b) divided Draba oreibata into two varieties: the Idaho endemic var. oreibata and the Nevada endemic var. serpentina. As indicated by I. A. Al-Shehbaz and M. D. Windham (2007), the two taxa are morphologically distinct, have different chromosome numbers, and are separated by over 480 kilometers. They are treated herein as separate species, and distinguished by characteristics discussed by Al-Shehbaz and Windham. Draba oreibata, in the strict sense, is known from Blaine, Butte, Clark, Custer, and Lemhi counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological and chromosomal data suggest that Draba spectabilis is an allopolyploid resulting from hybridization between D. abajoensis and D. malpighiacea. The characteristics distinguishing these three taxa were discussed in detail by I. A. Al-Shehbaz and M. D. Windham (2007). Chromosome counts attributed to D. spectabilis by Windham (2000) represent diploid D. abajoensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 325. | FNA vol. 7, p. 339. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. oxyloba, D. spectabilis var. bella, D. spectabilis var. oxyloba, D. spectabilis var. purpusii | |
Name authority | J. F. Macbride & Payson: Amer. J. Bot. 4: 257. (1917) | Greene: Pittonia 4: 19. (1899) |
Web links |