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limestone Draba

Macoun's Draba, Macoun's whitlow-grass

Habit Perennials; (cespitose); caudex branched (with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. Perennials; (cespitose); caudex simple or branched (with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose.
Stems

unbranched, 0.2–0.9 dm, glabrous throughout.

unbranched, (0.06–)0.1–0.4(–0.6) dm, usually pubescent, rarely glabrous, trichomes simple and stalked, 2–4-rayed, 0.15–0.9 mm.

Basal leaves

rosulate;

sessile;

blade oblong to narrowly oblanceolate, 0.2–1 cm × 0.6–2(–4) mm, margins entire, (ciliate, trichomes simple, 0.1–0.5 mm, apex obtuse), surfaces glabrous.

rosulate;

petiole (obscure), margin ciliate, (trichomes simple, 0.3–1 mm);

blade oblanceolate to obovate, (0.4–)0.6–1(–1.5) cm × (1–)2–4 mm, margins entire, surfaces usually pubescent abaxially, rarely glabrous, with short-stalked, 2–5-rayed trichomes, 0.1–0.5 mm, adaxially glabrous or pubescent with simple and short-stalked, 2-rayed trichomes.

Cauline leaves

0.

0.

Racemes

3–8-flowered, ebracteate, considerably elongated in fruit;

rachis slightly flexuous, glabrous.

3–10(–13)-flowered, ebracteate, (subumbellate), slightly elongated in fruit;

rachis not flexuous, usually pubescent as stem, rarely glabrous.

Flowers

sepals oblong, 1.7–2.5 mm, glabrous;

petals white, oblanceolate to obovate, 3.5–4.5 × 1.2–2 mm;

anthers ovate, 0.3–0.4 mm.

sepals ovate, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple and short-stalked, 2- or 3-rayed);

petals pale yellow, spatulate, 2.7–4 × 1–2 mm;

anthers ovate, 0.2–0.3 mm.

Fruiting pedicels

divaricate-ascending to ascending, straight, 4–13 mm, glabrous.

horizontal to divaricate (not expanded basally), curved upward, 1.5–4.5(–6) mm, abaxially pubescent as stem, adaxially usually glabrous, rarely throughout.

Fruits

oblong to narrowly so, slightly twisted, flattened, 5–9 × 1.5–3 mm;

valves glabrous;

ovules 8–16 per ovary;

style 0.3–0.9(–1.2) mm.

subglobose to ovoid or ellipsoid, plane, inflated at least basally, 4–8 × 2–4.5 mm;

valves glabrous;

ovules 10–14(–16) per ovary;

style 0.1–0.5 mm.

Seeds

oblong, 0.9–1.3 × 0.6–0.8 mm.

ovoid, 1–1.2 × 0.6–0.8 mm.

2n

= 32.

= 64.

Draba oreibata

Draba macounii

Phenology Flowering May–Jul. Flowering Jun–Aug.
Habitat Limestone cliffs, talus Rock outcrops, talus, tundra
Elevation 1800-2700 m (5900-8900 ft) 700-2800 m (2300-9200 ft)
Distribution
from FNA
ID
[BONAP county map]
from FNA
AK; MT; AB; BC; YT
[BONAP county map]
Discussion

R. C. Rollins (1993) and N. H. Holmgren (2005b) divided Draba oreibata into two varieties: the Idaho endemic var. oreibata and the Nevada endemic var. serpentina. As indicated by I. A. Al-Shehbaz and M. D. Windham (2007), the two taxa are morphologically distinct, have different chromosome numbers, and are separated by over 480 kilometers. They are treated herein as separate species, and distinguished by characteristics discussed by Al-Shehbaz and Windham. Draba oreibata, in the strict sense, is known from Blaine, Butte, Clark, Custer, and Lemhi counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba macounii is often confused with the circumpolar decaploid (2n = 80) D. alpina. Plants of the former have scapes (0.6–)1–4(–6) cm, fruiting pedicels divaricate to horizontal, gently curved, and 1.5–4.5(–6) mm, and petals pale yellow, spatulate, and 2.7–4 × 1–2 mm. By contrast, D. alpina has scapes (3–)5–17(–28) cm, fruiting pedicels divaricate-ascending to ascending, often straight, and (3–)4–14(–30) mm, and petals bright yellow, obovate, and 3.5–5 × 1.7–2.5 mm. R. C. Rollins (1993) indicated that the species occurs in Colorado, but we have not seen any material from that state.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 325. FNA vol. 7, p. 318.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Name authority J. F. Macbride & Payson: Amer. J. Bot. 4: 257. (1917) O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 97. (1927)
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