Draba novolympica |
Draba ramosissima |
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Mt. Olympic Draba, Olympic Draba, Olympic Mountains Draba, Payson's Draba, Payson's whitlow-grass, Trelease's Draba, Trelease's whitlow-grass |
branch Draba |
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Habit | Perennials; (cespitose, densely pulvinate); caudex branched (covered with persistent leaves, branches creeping, sometimes terminating in sterile rosettes); scapose. | Perennials; caudex branched (well-developed, with persistent leaf bases); not scapose. |
Stems | unbranched, 0.05–0.4 dm, densely pubescent throughout, trichomes simple, 0.4–0.8 mm, and stalked, 2–5-rayed, 0.1–0.5 mm. |
unbranched proximally, paniculately branched distally, 0.7–3.8(–5) dm, pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.1–0.6 mm, sometimes with simple ones, 0.4–0.8 mm. |
Basal leaves | (densely imbricate); rosulate; sessile; blade oblong to linear-oblanceolate, 0.2–0.8 cm × 0.5–1.5 mm, margins entire, (ciliate, trichomes simple and spurred, 0.3–1.2 mm), surfaces densely pubescent, abaxially with stalked, 2–12-rayed stellate trichomes, 0.1–0.6 mm, adaxially with simple and 2-rayed ones, 0.3–0.8 mm. |
rosulate; petiolate; petiole (to 2 cm), ciliate (trichomes simple, 0.3–0.5 mm); blade oblanceolate, (0.7–)1–5.5 cm × 2–8 mm, margins dentate, surfaces pubescent with short-stalked to subsessile, 3- or 4-rayed trichomes, 0.1–0.5 mm, adaxially sometimes also simple, 0.4–1.3 mm. |
Cauline leaves | 0. |
(5–)7–15(–21); sessile; blade ovate to oblong or elliptic to lanceolate, margins usually coarsely dentate or serrate, rarely subpinnatifid, surfaces pubescent as basal. |
Racemes | 2–12-flowered, ebracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
9–28-flowered, ebracteate, paniculate; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 1.5–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals bright yellow, oblanceolate to spatulate, 2–3.5(–4) × 1.5–2 mm; anthers oblong, 0.5–0.6 mm. |
sepals oblong, 2.5–3 mm, pubescent, (trichomes simple and subsessile, 2–4-rayed); petals white, spatulate, 4–6 × 1.5–2.5 mm; anthers oblong, 0.5–0.6 mm. |
Fruiting pedicels | divaricate-ascending, straight, 1–5 mm, pubescent, trichomes simple (0.3–0.9 mm) and stalked, 2–5-rayed (0.1–0.5 mm). |
horizontal to divaricate-ascending, straight, (4–)6–12(–15) mm, pubescent, trichomes 2–4-rayed. |
Fruits | often ovoid, plane, slightly inflated basally (symmetric), (2.5–)3–4(–5) × 1.5–3.5 mm; valves densely pubescent, trichomes 2–6-rayed, 0.05–0.4 mm, occasionally some simple; ovules 4–8(–12) per ovary; style 0.2–0.6(–0.8) mm. |
narrowly oblong to elliptic or oblong-lanceolate, often strongly twisted, flattened, 5–11 × 1.5–2.2 mm; valves pubescent, trichomes subsessile or sessile, 2–4-rayed, 0.07–0.3 mm; ovules 4–10(–12) per ovary; style (1)1.5–3(–4) mm, (sparsely pubescent). |
Seeds | oblong, 1.2–1.8 × 0.8–1.1 mm. |
oblong, 1.2–1.8 × 0.6–0.9 mm. |
2n | = 42. |
= 16. |
Draba novolympica |
Draba ramosissima |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Jun. |
Habitat | Alpine crests, open knolls, fellfields, talus, weathered shale, calcareous shale scree, rocky grounds and cliffs, subalpine conifer forests | Limestone cliffs and outcrops, shale barrens, rocky wooded areas |
Elevation | 1500-3700 m (4900-12100 ft) | 0-1100 m (0-3600 ft) |
Distribution |
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
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DC; KY; MD; NC; TN; VA; WV |
Discussion | Draba novolympica is the same taxon that C. L. Hitchcock (1941) and R. C. Rollins (1993) called D. paysonii var. treleasei, and G. A. Mulligan (2002) called D. paysonii. The two are amply distinct and should be recognized as separate species. Draba novolympica is easily distinguished from D. paysonii by having fruit valves pubescent with 2–6-rayed (occasionally some simple) trichomes 0.05–0.4 mm, sepals 1.5–2.5 mm, petals 2–3.5(–4) × 1.5–2 mm, fruits (2.5–)3–4(–5) × 1.5–3.5 mm, styles 0.2–0.6(–0.8) mm, and ovules 1.2–1.8 × 0.8–1.1 mm. By contrast, D. paysonii has fruit valves pubescent with simple and 2-rayed (some 4- or 5-rayed) trichomes (0.2–)0.4–1 mm, sepals 2.8–3.5 mm, petals (4–)5–6 × (1.5–)2–3 mm, fruits (5–)6–9 × (3–)3.5–5 mm, styles (0.6–)0.8–1.2 mm, and ovules 1.7–2.2 × 1–1.4 mm. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) indicated that Draba novolympica (as D. paysonii var. treleasei) occurs in Alaska and Yukon, but we have not seen any material from there, and it is likely that their records were based on misidentified plants. Previous reports of D. paysonii from Canada (e.g., G. A. Mulligan 1971b) pertain instead to D. novolympica. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 323. | FNA vol. 7, p. 332. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. barbata var. treleasei, D. paysonii var. treleasei | Alyssum dentatum, D. dentata, D. ramosissima var. glabrifolia |
Name authority | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 113. (1930) | Desvaux: J. Bot. Agric. 3: 186. (1815) |
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