Draba novolympica |
Draba oxycarpa |
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Mt. Olympic Draba, Olympic Draba, Olympic Mountains Draba, Payson's Draba, Payson's whitlow-grass, Trelease's Draba, Trelease's whitlow-grass |
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Habit | Perennials; (cespitose, densely pulvinate); caudex branched (covered with persistent leaves, branches creeping, sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (covered with persistent petiole remains); scapose. |
Stems | unbranched, 0.05–0.4 dm, densely pubescent throughout, trichomes simple, 0.4–0.8 mm, and stalked, 2–5-rayed, 0.1–0.5 mm. |
unbranched, 0.2–1.1 dm, pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.1–0.5 mm, (some rays spurred). |
Basal leaves | (densely imbricate); rosulate; sessile; blade oblong to linear-oblanceolate, 0.2–0.8 cm × 0.5–1.5 mm, margins entire, (ciliate, trichomes simple and spurred, 0.3–1.2 mm), surfaces densely pubescent, abaxially with stalked, 2–12-rayed stellate trichomes, 0.1–0.6 mm, adaxially with simple and 2-rayed ones, 0.3–0.8 mm. |
rosulate; petiolate; petiole base and margin ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to lanceolate, 0.8–2.2 cm × 2–6 mm, margins often entire, surfaces pubescent abaxially with stalked, 2–5-rayed, stellate trichomes, 0.1–0.4 mm, often some rays spurred and trichomes appearing subdendritic, rarely glabrate, or with simple and 2-rayed ones, (midvein obscure abaxially, not thickened), adaxially glabrous or sparsely pubescent with simple trichomes, sometimes with long-stalked, 2- or 3-rayed ones. |
Cauline leaves | 0. |
0. |
Racemes | 2–12-flowered, ebracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
(2–)4–13-flowered, ebracteate, (subcorymbose), not elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 1.5–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals bright yellow, oblanceolate to spatulate, 2–3.5(–4) × 1.5–2 mm; anthers oblong, 0.5–0.6 mm. |
sepals (grayish white), ovate to broadly so, 2.5–3 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals creamy white to yellow, broadly obovate, 3.5–5 × 2.5–3.5 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | divaricate-ascending, straight, 1–5 mm, pubescent, trichomes simple (0.3–0.9 mm) and stalked, 2–5-rayed (0.1–0.5 mm). |
subhorizontal to divaricate-ascending (not appressed to rachis), often slightly curved upward, 4–7(–10) mm, pubescent, trichomes simple (0.4–1 mm), and 2–4-rayed (0.1–0.4 mm). |
Fruits | often ovoid, plane, slightly inflated basally (symmetric), (2.5–)3–4(–5) × 1.5–3.5 mm; valves densely pubescent, trichomes 2–6-rayed, 0.05–0.4 mm, occasionally some simple; ovules 4–8(–12) per ovary; style 0.2–0.6(–0.8) mm. |
ovate to ovate-oblong, plane, flattened, 5–10 × 3–5 mm; valves puberulent along replum, trichomes simple, 0.1–0.3 mm; ovules (16–)18–28 per ovary; style 0.2–0.6 mm (stigma as wide as style). |
Seeds | oblong, 1.2–1.8 × 0.8–1.1 mm. |
(black), ovoid, 1–1.3 × 0.6–0.8 mm. |
2n | = 42. |
= 64. |
Draba novolympica |
Draba oxycarpa |
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Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Alpine crests, open knolls, fellfields, talus, weathered shale, calcareous shale scree, rocky grounds and cliffs, subalpine conifer forests | Rocky soils |
Elevation | 1500-3700 m (4900-12100 ft) | |
Distribution |
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
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Greenland; Europe (Norway [including Spitsbergen], Sweden); Atlantic Islands (Iceland) |
Discussion | Draba novolympica is the same taxon that C. L. Hitchcock (1941) and R. C. Rollins (1993) called D. paysonii var. treleasei, and G. A. Mulligan (2002) called D. paysonii. The two are amply distinct and should be recognized as separate species. Draba novolympica is easily distinguished from D. paysonii by having fruit valves pubescent with 2–6-rayed (occasionally some simple) trichomes 0.05–0.4 mm, sepals 1.5–2.5 mm, petals 2–3.5(–4) × 1.5–2 mm, fruits (2.5–)3–4(–5) × 1.5–3.5 mm, styles 0.2–0.6(–0.8) mm, and ovules 1.2–1.8 × 0.8–1.1 mm. By contrast, D. paysonii has fruit valves pubescent with simple and 2-rayed (some 4- or 5-rayed) trichomes (0.2–)0.4–1 mm, sepals 2.8–3.5 mm, petals (4–)5–6 × (1.5–)2–3 mm, fruits (5–)6–9 × (3–)3.5–5 mm, styles (0.6–)0.8–1.2 mm, and ovules 1.7–2.2 × 1–1.4 mm. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) indicated that Draba novolympica (as D. paysonii var. treleasei) occurs in Alaska and Yukon, but we have not seen any material from there, and it is likely that their records were based on misidentified plants. Previous reports of D. paysonii from Canada (e.g., G. A. Mulligan 1971b) pertain instead to D. novolympica. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba oxycarpa is an octoploid that most closely resembles the decaploid D. alpina. It is distinguished from that species by having creamy white to pale yellow (versus bright yellow) petals, gray-green (versus purplish tinged) sepals, petals 2.5–3.5 (versus 1.7–2.5) mm wide, stems pubescent proximally with primarily branched (versus primarily simple) trichomes, nearly black (versus pale brown) seeds, and ovate to ovate-oblong (versus elliptic) fruits 3–5 (versus 2–3) mm wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 323. | FNA vol. 7, p. 325. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. barbata var. treleasei, D. paysonii var. treleasei | D. alpina var. oxycarpa, D. gredinii |
Name authority | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 113. (1930) | Sommerfelt: Mag. Naturvidensk., n. s. 1: 240. (1833) |
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