Draba novolympica |
Draba californica |
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Mt. Olympic Draba, Olympic Draba, Olympic Mountains Draba, Payson's Draba, Payson's whitlow-grass, Trelease's Draba, Trelease's whitlow-grass |
California Draba |
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Habit | Perennials; (cespitose, densely pulvinate); caudex branched (covered with persistent leaves, branches creeping, sometimes terminating in sterile rosettes); scapose. | Perennials; (short-lived); caudex simple or branched; sometimes scapose. |
Stems | unbranched, 0.05–0.4 dm, densely pubescent throughout, trichomes simple, 0.4–0.8 mm, and stalked, 2–5-rayed, 0.1–0.5 mm. |
unbranched or branched (few), (0.2–)0.4–0.9(–1.2) dm, pubescent throughout, trichomes stellate, 4–8-rayed, 0.05–0.5 mm. |
Basal leaves | (densely imbricate); rosulate; sessile; blade oblong to linear-oblanceolate, 0.2–0.8 cm × 0.5–1.5 mm, margins entire, (ciliate, trichomes simple and spurred, 0.3–1.2 mm), surfaces densely pubescent, abaxially with stalked, 2–12-rayed stellate trichomes, 0.1–0.6 mm, adaxially with simple and 2-rayed ones, 0.3–0.8 mm. |
rosulate; shortly petiolate; petiole ciliate, margin not ciliate, (trichomes simple and 2-rayed, 0.3–1 mm); blade oblanceolate, 0.6–2 cm × 1.5–4(–6) mm, margins entire, surfaces pubescent with short-stalked, 4–8-rayed trichomes, 0.1–0.4 mm. |
Cauline leaves | 0. |
0–3; sessile; blade ovate to oblong, margins entire, surfaces pubescent as basal. |
Racemes | 2–12-flowered, ebracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
3–13-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 1.5–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals bright yellow, oblanceolate to spatulate, 2–3.5(–4) × 1.5–2 mm; anthers oblong, 0.5–0.6 mm. |
sepals (persistent until fruit maturity), oblong to ovate, 1.7–2.5 mm, pubescent, (trichomes short-stalked, 2–5-rayed); petals white or creamy white, spatulate to oblanceolate, 2.2–3 × 0.8–1 mm; anthers ovate, 0.2–0.35 mm. |
Fruiting pedicels | divaricate-ascending, straight, 1–5 mm, pubescent, trichomes simple (0.3–0.9 mm) and stalked, 2–5-rayed (0.1–0.5 mm). |
ascending to divaricate-ascending, straight, (2–)4–9(–12) mm, pubescent, trichomes short-stalked, 4–8-rayed, (0.1–0.4 mm). |
Fruits | often ovoid, plane, slightly inflated basally (symmetric), (2.5–)3–4(–5) × 1.5–3.5 mm; valves densely pubescent, trichomes 2–6-rayed, 0.05–0.4 mm, occasionally some simple; ovules 4–8(–12) per ovary; style 0.2–0.6(–0.8) mm. |
lanceolate to oblong, plane, slightly flattened, (5–)6–9(–11) × 1.8–2.5 mm; valves usually pubescent, rarely glabrous, trichomes simple and short-stalked, 2- or 3-rayed, 0.05–0.25 mm; ovules 22–32 per ovary; style 0.1–0.3(–0.4) mm. |
Seeds | oblong, 1.2–1.8 × 0.8–1.1 mm. |
oblong, 0.8–1 × 0.5–0.6 mm. |
2n | = 42. |
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Draba novolympica |
Draba californica |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Alpine crests, open knolls, fellfields, talus, weathered shale, calcareous shale scree, rocky grounds and cliffs, subalpine conifer forests | Grassy meadows, alpine areas, fellfields |
Elevation | 1500-3700 m (4900-12100 ft) | 3200-4000 m (10500-13100 ft) |
Distribution |
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
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CA; NV |
Discussion | Draba novolympica is the same taxon that C. L. Hitchcock (1941) and R. C. Rollins (1993) called D. paysonii var. treleasei, and G. A. Mulligan (2002) called D. paysonii. The two are amply distinct and should be recognized as separate species. Draba novolympica is easily distinguished from D. paysonii by having fruit valves pubescent with 2–6-rayed (occasionally some simple) trichomes 0.05–0.4 mm, sepals 1.5–2.5 mm, petals 2–3.5(–4) × 1.5–2 mm, fruits (2.5–)3–4(–5) × 1.5–3.5 mm, styles 0.2–0.6(–0.8) mm, and ovules 1.2–1.8 × 0.8–1.1 mm. By contrast, D. paysonii has fruit valves pubescent with simple and 2-rayed (some 4- or 5-rayed) trichomes (0.2–)0.4–1 mm, sepals 2.8–3.5 mm, petals (4–)5–6 × (1.5–)2–3 mm, fruits (5–)6–9 × (3–)3.5–5 mm, styles (0.6–)0.8–1.2 mm, and ovules 1.7–2.2 × 1–1.4 mm. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) indicated that Draba novolympica (as D. paysonii var. treleasei) occurs in Alaska and Yukon, but we have not seen any material from there, and it is likely that their records were based on misidentified plants. Previous reports of D. paysonii from Canada (e.g., G. A. Mulligan 1971b) pertain instead to D. novolympica. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although originally treated as a variety of Draba cuneifolia, D. californica is but distantly related to that species. Instead, ongoing studies (L. Allphin and M. D. Windham, unpubl.) suggest that it may be an allopolyploid resulting from hybridization between D. albertina and D. breweri. The species is restricted to the White Mountains in Mono County, California, and adjacent Esmeralda County, Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 323. | FNA vol. 7, p. 298. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. barbata var. treleasei, D. paysonii var. treleasei | D. cuneifolia var. californica |
Name authority | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 113. (1930) | (Jepson) Rollins & R. A. Price: Aliso 12: 19. (1988) |
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