Draba norvegica |
Draba exunguiculata |
|
---|---|---|
Norwegian Draba |
clawless Draba |
|
Habit | Perennials; (cespitose); caudex branched (sometimes with persistent leaf remains); often scapose. | Perennials; (densely cespitose); caudex branched (covered with persistent, somewhat thickened, dry petioles); not scapose. |
Stems | branched or unbranched, (0.2–)0.4–1.4(–2) dm, pubescent throughout, trichomes simple and 2–4-rayed, 0.2–0.5 mm. |
unbranched, 0.15–0.7 dm, glabrous throughout or sparsely pubescent, trichomes simple, 0.3–0.9 mm. |
Basal leaves | rosulate; petiolate; petiole (0–0.5 cm), ciliate throughout; blade oblanceolate to narrowly obovate or narrowly lanceolate, 0.4–2.1 cm × 1.5–7 mm, margins with 1–3 teeth on each side, (ciliate, trichomes simple and 2-rayed, 0.2–0.8 mm), surfaces pubescent abaxially with stalked, (2–)4(–6)-rayed trichomes, 0.1–0.4 mm, adaxially glabrous or pubescent, with simple and stalked, 2-rayed trichomes, 0.1–0.5 mm. |
rosulate; petiolate; petiole ciliate, (trichomes simple, 0.3–1.1 mm); blade linear to linear-oblanceolate, (0.5–)0.8–2(–2.5) cm × 1–3 mm, margins entire, (pubescent as petiole), surfaces abaxially sparsely pubescent with simple and stalked, 2- or 3-rayed trichomes, 0.1–0.4 mm, adaxially glabrous or subapically sparsely pubescent with simple trichomes. |
Cauline leaves | 0–3(–5); sessile; blade ovate to oblong, margins often dentate, (ciliate). |
1–4; sessile; blade linear to linear-oblanceolate, margins entire, surfaces pubescent as basal. |
Racemes | 5–23-flowered, ebracteate or proximalmost 1(–3) flowers bracteate, considerably elongated in fruit; rachis slightly or not flexuous, pubescent as stem. |
4–13(–20)-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent, trichomes simple, to 1 mm. |
Flowers | sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals white, spatulate, 2.5–4 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 2–2.5 mm, pubescent, (trichomes simple); petals (erect), yellow, obovate, 2–2.5(–3) × 1.5–2 mm, (not clawed); anthers oblong, 0.6–0.9 mm. |
Fruiting pedicels | suberect to ascending (often appressed to rachis), straight, (1.5–)2.5–5(–9) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending, straight or slightly curved upward, 1–6 mm, usually glabrous, rarely with few, simple trichomes. |
Fruits | (sometimes subappressed to rachis), oblong to lanceolate-elliptic, plane, flattened, 4–8 × 2–3 mm; valves glabrous or pubescent, trichomes simple and short-stalked, 2-rayed, 0.05–0.25 mm; ovules 18–26 per ovary; style (0.01–)0.1–0.4(–0.5) mm, (glabrous, stigma about as wide as style). |
lanceolate to oblong-lanceolate, plane, flattened, 5–13 × 1.5–3 mm; valves glabrous; ovules 20–24 per ovary; style 0.3–1 mm. |
Seeds | oblong, 0.9–1.2 × 0.5–0.7 mm. |
ovoid, 1–1.3 × 0.6–0.8 mm. |
2n | = 48. |
= 56 ± 5. |
Draba norvegica |
Draba exunguiculata |
|
Phenology | Flowering Jun–Aug. | Flowering Jul–Aug. |
Habitat | Rock outcrops and sea cliffs, talus, gravelly and sandy terraces, moist bluffs, turfy limestone shores | Open knolls, talus, gravelly alpine slopes, rocky alpine tundra |
Elevation | 0-500 m (0-1600 ft) | 3600-4300 m (11800-14100 ft) |
Distribution |
NL; NS; NT; QC; Greenland; Europe (Finland, Norway, n Russia, Scotland, Sweden); Atlantic Islands (Iceland) |
CO |
Discussion | Draba norvegica is a highly variable hexaploid taxon and, as delimited herein, it probably represents two taxa with the same distribution and same chromosome number. It is related to hexaploid D. arctogena, from which it is distinguished by characteristics listed under 8. D. arctogena. Draba norvegica was divided by O. E. Schulz (1927) into eight varieties, whereas R. C. Rollins (1993) recognized just two. In the latter treatment, var. clivicola is distinguished by having predominantly branched, appressed trichomes on stems proximally, lanceolate, glabrous fruits, and slender, erect-appressed pedicels. By contrast, var. norvegica is said to have predominantly simple, spreading trichomes on stems proximally, narrowly ovate to elliptical, glabrous or pubescent fruits, and stout, divaricately ascending pedicels. These differences are extremes of a continuum, and we agree with G. A. Mulligan (1970) in reducing D. clivicola to synonymy of D. norvegica. Rollins indicated that D. norvegica grows in Minnesota, but we have not seen any material from that state. Draba norvegica is found in Europe in arctic and subarctic areas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba exunguiculata occupies alpine areas of Clear Creek, El Paso, Grand, and Summit counties in central Colorado. Draba grayana is found in the same general area; both species are apomicts that occasionally grow sympatrically. Draba exunguiculata is easily distinguished from D. grayana by having stems and fruiting pedicels glabrous or sparsely pubescent with simple trichomes (versus moderately to densely pubescent with simple and 2- or 3-rayed trichomes), and by having erect, non-clawed petals, 2–3 mm and slightly longer than sepals (versus flared, clawed petals 3–4.5 mm and nearly twice as long as sepals). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 322. | FNA vol. 7, p. 306. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. clivicola, D. hirta var. norvegica, D. norvegica var. clivicola, D. norvegica var. hebecarpa, D. rupestris, D. scandinavica var. hebecarpa, D. trichella | D. chrysantha var. exunguiculata, D. chrysantha |
Name authority | Gunnerus: Fl. Norveg. 2: 106. (1772) | (O. E. Schulz) C. L. Hitchcock: Revis. Drabas W. N. Amer., 46. (1941) |
Web links |