Draba nivalis |
Draba streptobrachia |
|
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snow Draba, snow whitlow-grass, yellow arctic Draba |
alpine tundra Draba |
|
Habit | Perennials; (cespitose, usually forming mats); caudex branched (covered with persistent leaf bases); usually scapose. | Perennials; caudex branched (often with persistent leaf remains, branches not creeping); not scapose. |
Stems | unbranched, 0.2–0.8(–1.2) dm, pubescent throughout, trichomes minutely stalked, 8–15-rayed, stellate, (non-crisped), 0.03–0.15 mm. |
unbranched, (0.1–)0.2–1(–1.3) dm, pubescent throughout, trichomes subsessile (often crisped), 3–5-rayed, stellate, 0.03–0.25(–0.4) mm, (rays sometimes forked). |
Basal leaves | rosulate; petiole (obsolete), ciliate throughout; blade oblanceolate to obovate, 0.2–0.9(–1.5) cm × 1–3(–5) mm, margins entire, (not ciliate), surfaces usually pubescent, rarely glabrescent, with short-stalked, 8–15-rayed, stellate, (non-crisped) trichomes, 0.05–0.15 mm. |
rosulate; petiole (obscure), usually not ciliate, rarely sparsely pubescent, (trichomes simple, to 0.6 mm); blade oblanceolate to linear-oblanceolate, (0.4–)0.5–3(–4) cm × 1–5 mm, margins entire, surfaces pubescent with short-stalked (crisped), 3–8-rayed trichomes, 0.05–0.4 mm. |
Cauline leaves | 0 or 1; sessile; blade ovate or oblong, margins entire, surfaces pubescent as basal. |
(1 or) 2–4 (or 5); sessile; blade oblong to ovate or linear, margins entire, surfaces pubescent as basal. |
Racemes | 3–9(–11)-flowered, ebracteate, elongated in fruit; rachis often slightly flexuous, pubescent as stem. |
4–10(–18)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and short-stalked, 2–5-rayed); petals white, spatulate to oblanceolate, 2–3.5 × 0.8–1.4 mm; anthers ovate, 0.2–0.3 mm. |
sepals ovate, 2–3 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals yellow, spatulate, 3–5 × 1.5–3 mm, (clawed); anthers ovate, 0.25–0.4 mm. |
Fruiting pedicels | usually ascending, rarely divaricate, straight, 1–4.5(–8) mm, pubescent as stem. |
ascending, usually straight, rarely curved upward, (2–)3–8(–12) mm, pubescent as stem. |
Fruits | elliptic to narrowly oblong-elliptic, twisted or not, flattened, 3.5–9 × 1.5–2.2 mm; valves usually glabrous, rarely with 3–6-rayed trichomes on replum; ovules 12–24(–28) per ovary; style 0.1–0.4 mm. |
(not appressed to rachis), ovate to elliptic or lanceolate, slightly twisted or plane, flattened, (3–)5–10 × 2–4 mm; valves often pubescent, occasionally glabrous, trichomes simple and minutely stalked, 2–4-rayed, 0.03–0.25 mm; ovules 10–16(–18) per ovary; style 0.3–0.8(–1.2) mm. |
Seeds | ovoid, 0.6–1 × 0.5–0.6 mm. |
oblong, 1–1.6 × 0.6–1 mm. |
2n | = 16. |
= ca. 64. |
Draba nivalis |
Draba streptobrachia |
|
Phenology | Flowering Jun–Aug. | Flowering Jul–Aug. |
Habitat | Rock outcrops and fellfields, meadows, open tundra, stream banks, gravelly beaches, roadsides | Alpine tundra, scree, ridges and alpine slopes, turf, fellfields, talus slopes, crevices in rock ledges, loose soils |
Elevation | 0-2500 m (0-8200 ft) | 3200-4000 m (10500-13100 ft) |
Distribution |
AK; AB; BC; MB; NF; NT; NU; ON; QC; YT; Greenland; Europe (Finland, Norway [including Svalbard], n, w Sweden); e Asia (Russian Far East, n Siberia); Atlantic Islands (Iceland); circumpolar |
CO
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Discussion | Draba nivalis is most often confused with D. lonchocarpa, and C. L. Hitchcock (1941) reduced the latter to a variety (var. elongata) of D. nivalis. A critical comparison reveals that there are enough differences to warrant their recognition as distinct species. Draba nivalis is distinguished by having non-ciliate leaf bases, basal leaf blades pubescent with trichomes 0.05–0.15 mm, stems and pedicels that are always pubescent, and fruits that are elliptic to narrowly elliptic-oblong, plane, and 3.5–9 mm. By contrast, D. lonchocarpa has ciliate leaf bases, basal leaf blades pubescent with trichomes 0.15–0.5 mm, stems and pedicels that are glabrous or sparsely pubescent, and fruits that are linear to lanceolate or oblong, slightly twisted or plane, and 6–15(–18) mm. Except for Alaska, D. nivalis does not grow in the continental United States, whereas D. lonchocarpa grows in nearly all Mountain and Pacific states. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As indicated by Price, Draba streptobrachia is an apomict, yielding abundant, well-developed seed despite producing only abortive pollen. Morphological studies (M. D. Windham, unpubl.) suggest that the species may be an allopolyploid containing a genome from D. crassa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 322. | FNA vol. 7, p. 341. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. caesia | D. chrysantha, D. spectabilis var. dasycarpa |
Name authority | Liljeblad: Utkast Sv. Fl., 236. (1792) | R. A. Price: Brittonia 32: 168. (1980) |
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