Draba nivalis |
Draba stenopetala |
|
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snow Draba, snow whitlow-grass, yellow arctic Draba |
Anadyr Draba, star-flower Draba, star-flower whitlow-grass |
|
Habit | Perennials; (cespitose, usually forming mats); caudex branched (covered with persistent leaf bases); usually scapose. | Perennials; (pulvinate); caudex branched (covered with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, 0.2–0.8(–1.2) dm, pubescent throughout, trichomes minutely stalked, 8–15-rayed, stellate, (non-crisped), 0.03–0.15 mm. |
unbranched, (0.02–)0.07–0.2(–0.3) dm, pubescent throughout, trichomes simple and short-stalked, 2–5-rayed, 0.1–0.5 mm, (sometimes simple ones very sparse). |
Basal leaves | rosulate; petiole (obsolete), ciliate throughout; blade oblanceolate to obovate, 0.2–0.9(–1.5) cm × 1–3(–5) mm, margins entire, (not ciliate), surfaces usually pubescent, rarely glabrescent, with short-stalked, 8–15-rayed, stellate, (non-crisped) trichomes, 0.05–0.15 mm. |
(densely imbricate); rosulate; petiolate; petiole base and margin ciliate, (trichomes simple and 2-rayed, 0.2–0.5 mm); blade obovate to oblong-spatulate, 0.2–0.6(–0.8) cm × 1–2(–3) mm, margins entire, surfaces often pubescent with simple and stalked, 2–5-rayed trichomes, 0.2–0.9 mm, sometimes glabrous and trichomes on margins only. |
Cauline leaves | 0 or 1; sessile; blade ovate or oblong, margins entire, surfaces pubescent as basal. |
0. |
Racemes | 3–9(–11)-flowered, ebracteate, elongated in fruit; rachis often slightly flexuous, pubescent as stem. |
2–5-flowered, ebracteate, not or slightly elongated in fruit; rachis slightly flexuous, pubescent as stem. |
Flowers | sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and short-stalked, 2–5-rayed); petals white, spatulate to oblanceolate, 2–3.5 × 0.8–1.4 mm; anthers ovate, 0.2–0.3 mm. |
sepals (spreading or reflexed), oblong, 2–3.5 mm, pubescent, (trichomes simple and 2-rayed); petals yellow or purple, linear, 2.5–5 × 0.3–0.7 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | usually ascending, rarely divaricate, straight, 1–4.5(–8) mm, pubescent as stem. |
divaricate, straight, 2–5(–7) mm, pubescent as stem. |
Fruits | elliptic to narrowly oblong-elliptic, twisted or not, flattened, 3.5–9 × 1.5–2.2 mm; valves usually glabrous, rarely with 3–6-rayed trichomes on replum; ovules 12–24(–28) per ovary; style 0.1–0.4 mm. |
suborbicular, plane, inflated basally, flattened distally, 3–5 × 3–4 mm; valves glabrous or puberulent, trichomes simple, 0.02–0.3 mm; ovules 4–10 per ovary; style 0.2–0.6 mm. |
Seeds | ovoid, 0.6–1 × 0.5–0.6 mm. |
ovoid, 1–1.3 × 0.8–1 mm. |
2n | = 16. |
= 24, 64. |
Draba nivalis |
Draba stenopetala |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Rock outcrops and fellfields, meadows, open tundra, stream banks, gravelly beaches, roadsides | Rock outcrops, talus, rocky ridges, alpine tundra |
Elevation | 0-2500 m (0-8200 ft) | 100-1900 m (300-6200 ft) |
Distribution |
AK; AB; BC; MB; NF; NT; NU; ON; QC; YT; Greenland; Europe (Finland, Norway [including Svalbard], n, w Sweden); e Asia (Russian Far East, n Siberia); Atlantic Islands (Iceland); circumpolar |
AK; YT; e Asia (Russian Far East) |
Discussion | Draba nivalis is most often confused with D. lonchocarpa, and C. L. Hitchcock (1941) reduced the latter to a variety (var. elongata) of D. nivalis. A critical comparison reveals that there are enough differences to warrant their recognition as distinct species. Draba nivalis is distinguished by having non-ciliate leaf bases, basal leaf blades pubescent with trichomes 0.05–0.15 mm, stems and pedicels that are always pubescent, and fruits that are elliptic to narrowly elliptic-oblong, plane, and 3.5–9 mm. By contrast, D. lonchocarpa has ciliate leaf bases, basal leaf blades pubescent with trichomes 0.15–0.5 mm, stems and pedicels that are glabrous or sparsely pubescent, and fruits that are linear to lanceolate or oblong, slightly twisted or plane, and 6–15(–18) mm. Except for Alaska, D. nivalis does not grow in the continental United States, whereas D. lonchocarpa grows in nearly all Mountain and Pacific states. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Different chromosome counts for Draba stenopetala were reported as 2n = 24 from North America and 2n = 64 from the Russian Far East (S. I. Warwick and I. A. Al-Shehbaz 2006). It is unlikely that a single species is involved, and further work is needed to verify the counts from North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 322. | FNA vol. 7, p. 341. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. caesia | D. stenopetala var. purpurea |
Name authority | Liljeblad: Utkast Sv. Fl., 236. (1792) | Trautvetter: Trudy Imp. S.-Petersburgsk. Bot. Sada 6: 11. (1879) |
Web links |