Draba monoensis |
Draba spectabilis |
|
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Mono Draba, White Mountains Draba |
showy Draba, showy whitlowgrass |
|
Habit | Perennials; (usually cespitose); caudex simple or branched (poorly developed, with persistent leaf remains); sometimes scapose. | Perennials; caudex simple or branched (sometimes with persistent leaf bases); not scapose. |
Stems | unbranched, (0.05–)0.1–0.4 dm, usually pubescent throughout, rarely sparsely pubescent or glabrous distally, trichomes simple and 2-rayed, (often crisped), 0.1–0.6 mm. |
unbranched, (0.7–)1.1–3.7(–5.3) dm, pubescent throughout (sparsely so distally), trichomes simple, malpighiaceous, or sessile and 4-rayed, 0.15–0.5 mm, (2 rays parallel to stem axis longer). |
Basal leaves | rosulate; subsessile or shortly petiolate; petiole margin ciliate or not; blade narrowly oblanceolate, (0.3–)0.5–1.6(–2) cm × (1–)1.5–3(–4) mm, margins usually entire, rarely subapically denticulate, (ciliate or not), surfaces pubescent with simple and stalked, 2-rayed trichomes, 0.1–0.7 mm, (midvein obscure abaxially). |
rosulate; petiolate; petiole sometimes ciliate, (trichomes simple); blade oblanceolate to spatulate or obovate, (1–)1.5–4.4(–6.3) cm × (3–)5–12(–15) mm, margins entire or dentate, surfaces pubescent with sessile, often 4-rayed trichomes, 0.1–0.6 mm (longest rays parallel to midvein, lateral rays sometimes reduced to tiny spurs), sometimes malpighiaceous trichomes present. |
Cauline leaves | 0–2 (or 3); sessile; blade ovate to oblong, margins entire, surfaces pubescent as basal. |
4–12(–17); sessile; blade broadly ovate to lanceolate or oblong, margins entire or dentate, (ciliate or not), surfaces pubescent as basal or adaxially also with simple trichomes. |
Racemes | (3–)6–13(–17)-flowered, ebracteate or proximalmost 1 (or 2) flowers bracteate, not or slightly elongated in fruit; rachis not flexuous, usually pubescent as stem, rarely glabrous. |
(10–)16–49(–61)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals (persistent), oblong, 1–1.5 mm, subapically sparsely pubescent, (trichomes simple); petals white, spatulate, 1.5–2 × 0.5–0.6 mm; anthers ovate, 0.15–0.2 mm. |
sepals ovate, (2.2–)2.5–4 mm, glabrous or pubescent, (trichomes simple and malpighiaceous); petals yellow, oblanceolate, 4–6.5 × 1–2.5 mm; anthers oblong, 0.5–0.7 mm. |
Fruiting pedicels | divaricate-ascending, straight, 1–2.5(–4) mm, usually pubescent as stem, rarely glabrous. |
horizontal to divaricate-ascending, straight or curved upward, (5–)7–20(–26) mm, glabrous or pubescent as rachis abaxially. |
Fruits | ovoid to subellipsoid, plane, slightly inflated basally, (2–)3–5 × (1.2–)1.5–2.5 mm; valves usually puberulent, rarely glabrous, trichomes simple, 0.05–0.2 mm; ovules 12–20 per ovary; style 0.1–0.2 mm. |
lanceolate to oblong or elliptic, plane, flattened, (6–)7–13 × 2–3.5 mm; valves glabrous or puberulent, trichomes simple and sessile, 2-rayed, 0.03–0.3 mm; ovules 12–24 per ovary; style (0.5–)1–2.7 mm. |
Seeds | ovoid, 0.6–0.8 × 0.4–0.6 mm. |
ovoid, 1–1.4 × 0.7–0.9 mm. |
2n | = 40. |
|
Draba monoensis |
Draba spectabilis |
|
Phenology | Flowering Jul–Aug. | Flowering Jun–Aug. |
Habitat | Gravelly alpine meadows | Talus, rocky hillsides, meadows in open conifer forests, aspen groves, and alpine communities |
Elevation | 3600-4000 m (11800-13100 ft) | 2000-3900 m (6600-12800 ft) |
Distribution |
CA |
CO; NM; WY
|
Discussion | Of conservation concern. Draba monoensis appears to be polyploid (M. D. Windham, unpubl.), and its morphological similarity to D. fladnizensis (R. C. Rollins 1993) may indicate that the latter is one of its progenitors. The species is known from the White Mountains of Mono County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological and chromosomal data suggest that Draba spectabilis is an allopolyploid resulting from hybridization between D. abajoensis and D. malpighiacea. The characteristics distinguishing these three taxa were discussed in detail by I. A. Al-Shehbaz and M. D. Windham (2007). Chromosome counts attributed to D. spectabilis by Windham (2000) represent diploid D. abajoensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 320. | FNA vol. 7, p. 339. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. oxyloba, D. spectabilis var. bella, D. spectabilis var. oxyloba, D. spectabilis var. purpusii | |
Name authority | Rollins & R. A. Price: Aliso 12: 22, figs. 1f–j, 3. (1988) | Greene: Pittonia 4: 19. (1899) |
Web links |