Draba monoensis |
Draba grandis |
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Mono Draba, White Mountains Draba |
north Pacific Draba |
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Habit | Perennials; (usually cespitose); caudex simple or branched (poorly developed, with persistent leaf remains); sometimes scapose. | Perennials; caudex simple or branched (fleshy, with persistent leaf bases); not scapose. |
Stems | unbranched, (0.05–)0.1–0.4 dm, usually pubescent throughout, rarely sparsely pubescent or glabrous distally, trichomes simple and 2-rayed, (often crisped), 0.1–0.6 mm. |
(decumbent), unbranched, (0.2–)0.5–2.7(–3.7) dm, pubescent throughout, trichomes 2–4-rayed, 0.05–0.2 mm. |
Basal leaves | rosulate; subsessile or shortly petiolate; petiole margin ciliate or not; blade narrowly oblanceolate, (0.3–)0.5–1.6(–2) cm × (1–)1.5–3(–4) mm, margins usually entire, rarely subapically denticulate, (ciliate or not), surfaces pubescent with simple and stalked, 2-rayed trichomes, 0.1–0.7 mm, (midvein obscure abaxially). |
rosulate; long-petiolate; petiole (winged, (1–)4–15 cm), often not ciliate (or ciliate to blade apex, trichomes simple, 0.3–0.8 mm); blade (somewhat fleshy), oblanceolate to spatulate or obovate, (1–)2–11(–17) cm × (5–)8–30(–45) mm, margins often coarsely dentate, (pubescent as petiole), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.25 mm, adaxially similar, or also with simple and long-stalked, 2-rayed trichomes, to 0.8 mm, rarely glabrescent, with mostly simple and 2-rayed trichomes. |
Cauline leaves | 0–2 (or 3); sessile; blade ovate to oblong, margins entire, surfaces pubescent as basal. |
2–12(–16); sessile or petiolate; blade oblanceolate to obovate, margins dentate or entire, surfaces pubescent as basal. |
Racemes | (3–)6–13(–17)-flowered, ebracteate or proximalmost 1 (or 2) flowers bracteate, not or slightly elongated in fruit; rachis not flexuous, usually pubescent as stem, rarely glabrous. |
5–26(–32)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals (persistent), oblong, 1–1.5 mm, subapically sparsely pubescent, (trichomes simple); petals white, spatulate, 1.5–2 × 0.5–0.6 mm; anthers ovate, 0.15–0.2 mm. |
sepals broadly ovate, 3–4 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, oblanceolate to spatulate, 4.5–7 × 1.8–3 mm; anthers oblong, 0.7–1 mm. |
Fruiting pedicels | divaricate-ascending, straight, 1–2.5(–4) mm, usually pubescent as stem, rarely glabrous. |
horizontal to divaricate-ascending or ascending, usually straight, rarely curved upward, (5–)10–22(–27) mm, pubescent, trichomes 2–4-rayed (0.05–0.3 mm), sometimes with simple and spurred ones. |
Fruits | ovoid to subellipsoid, plane, slightly inflated basally, (2–)3–5 × (1.2–)1.5–2.5 mm; valves usually puberulent, rarely glabrous, trichomes simple, 0.05–0.2 mm; ovules 12–20 per ovary; style 0.1–0.2 mm. |
oblong to lanceolate, or ovate to suborbicular, slightly twisted or plane, flattened, (6–)10–20(–25) × 4–7(–9) mm; valves glabrous; ovules 24–52 per ovary; style (0.2–)0.4–1.6(–2) mm. |
Seeds | ovoid, 0.6–0.8 × 0.4–0.6 mm. |
ovoid, 1.4–2 × 0.8–1.2 mm. |
2n | = 36. |
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Draba monoensis |
Draba grandis |
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Phenology | Flowering Jul–Aug. | Flowering May–Jul. |
Habitat | Gravelly alpine meadows | Rocky bluffs above salt-water beaches, loamy seaside banks, sea bird rookeries, coastal herbaceous tundra and sandy blowouts |
Elevation | 3600-4000 m (11800-13100 ft) | 0-50(-200) m (0-200(-700) ft) |
Distribution |
CA |
AK; BC; e Asia (Kuril and Ratmanov islands, Russian Far East) |
Discussion | Of conservation concern. Draba monoensis appears to be polyploid (M. D. Windham, unpubl.), and its morphological similarity to D. fladnizensis (R. C. Rollins 1993) may indicate that the latter is one of its progenitors. The species is known from the White Mountains of Mono County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Almost all recent North American authors have used the name Draba hyperborea for this species. A. N. Berkutenko (1995) clearly showed that the type of that name belongs to an entirely different species that she placed in the genus Schivereckia Andrzejowski ex de Candolle. Draba grandis thus becomes the correct name for the North American taxon. Except for its fleshy leaves, Nesodraba is indistinguishable morphologically and molecularly from other species of Draba. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 320. | FNA vol. 7, p. 309. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Cochlearia siliquosa, Cochlearia spathulata, D. greenei, D. hatchiae, D. hyperborea var. spathulata, D. spathulata, Nesodraba grandis, Nesodraba megalocarpa, Nesodraba siliquosa | |
Name authority | Rollins & R. A. Price: Aliso 12: 22, figs. 1f–j, 3. (1988) | Langsdorff ex de Candolle: Syst. Nat. 2: 355. (1821) |
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