Draba juvenilis |
Draba carnosula |
|
---|---|---|
long-stalk Draba, long-stalk whitlow-grass |
Mount Eddy Draba, Mt. Eddy Draba |
|
Habit | Perennials; (not pulvinate); caudex branched (branches slightly loose, with persistent leaf remains, sometimes terminating in sterile rosettes); often scapose. | Perennials; (loosely cespitose); caudex branched (somewhat surculose, with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.3–)0.8–2.3(–3) dm, pubescent proximally, glabrous or sparsely pubescent distally, trichomes simple and 2–4-rayed, 0.07–0.4 mm. |
unbranched, 0.3–1.2 dm, glabrous throughout. |
Basal leaves | (loosely) rosulate; petiolate; petiole base ciliate, margin not ciliate, (midvein obscure, trichomes simple and 2-rayed, 0.2–0.7 mm); blade oblanceolate to lanceolate, (0.3–)0.6–2.5(–3.5) cm × (1–)2–7(–10) mm, margins entire or denticulate, surfaces pubescent, abaxially with stalked, cruciform trichomes, 0.2–0.5 mm, adaxially with cruciform and/or simple and 2-rayed ones. |
rosulate; petiolate; petiole (persistent, midvein prominent), ciliate, (trichomes sparse, short-stalked, 2–4-rayed, sometimes with simple ones, 0.2–0.5 mm); blade (somewhat fleshy), oblanceolate or spatulate to obovate, 0.3–1.5 cm × 1.5–5 mm, margins entire, (pubescent as petiole), surfaces glabrous. |
Cauline leaves | 0–2 (or 3); sessile; blade elliptic to ovate or lanceolate, margins usually entire, rarely denticulate, surfaces pubescent as basal adaxially or trichomes predominantly simple. |
0. |
Racemes | (2–)4–13(–18)-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous or sparsely pubescent as stem. |
2–6-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals ovate, 2.2–3 mm, glabrous or pubescent subapically, (trichomes simple and short-stalked, 2-rayed); petals pale yellow to creamy white, spatulate, 3–5 × 1.5–2.5 mm; anthers ovate, 0.4–0.5 mm. |
sepals ovate, 3–4 mm, glabrous; petals yellow, oblanceolate, 5–7 × 1.3–2 mm; anthers oblong, 0.7–0.9 mm. |
Fruiting pedicels | divaricate-ascending or ascending, straight or often curved upward (not expanded basally), (3–)5–17(–22) mm, usually glabrous, rarely pubescent, trichomes simple and stalked, 2–4-rayed. |
divaricate-ascending, straight, 3–8 mm, usually glabrous, rarely trichomes simple. |
Fruits | elliptic to oblong or linear-lanceolate, plane (not curved), flattened, 5–11(14) × 2–3 mm; valves usually glabrous, rarely margin pubescent, trichomes simple; ovules 16–30 per ovary; style 0.2–0.7 mm. |
lanceolate to elliptic-lanceolate, plane, strongly flattened, 10–23 × 4–6 mm; valves (each with distinct midvein), glabrous; ovules 8–12 per ovary; style 2–3 mm. |
Seeds | oblong, 0.7–1.2(–1.5) × 0.4–0.7(–0.8) mm. |
(broadly winged), orbicular, 3–4.5 in diam.; (wing 1–1.5 mm wide). |
2n | = 64. |
|
Draba juvenilis |
Draba carnosula |
|
Phenology | Flowering Jun–Jul. | Flowering Jun–Jul. |
Habitat | Rock outcrops, talus, gravelly beaches and stream banks, meadows, tundra | Open rocky hillsides |
Elevation | 0-2700 m (0-8900 ft) | 2800-3100 m (9200-10200 ft) |
Distribution |
AK; AB; BC; NT; YT; e Asia (Russian Far East, n Siberia) |
CA |
Discussion | Although Draba kananaskis and D. longipes have often been treated as distinct species, we find no basis for maintaining them. Both exhibit variations in petal color (white to pale yellow) and leaf trichomes (short-stalked to sessile) characteristic of D. juvenilis from the Russian Far East. Perhaps most importantly, all three are octoploids (2n = 64) with x = 8. We conclude that D. kananaskis is nothing more than a minor variant of D. juvenilis and it is treated herein, for the first time, as a synonym of that species. Draba juvenilis is occasionally confused with D. borealis, which also has stalked, cruciform trichomes with unbranched rays. Typical D. juvenilis is easily distinguished from that species by having narrower (2–3 mm) fruits that are glabrous (rarely pubescent) and untwisted, and 0–2 (or 3)-leaved stems. By contrast, D. borealis has wider (2.5–4.5 mm) fruits that are usually pubescent and/or twisted (rarely neither) and (2 or) 3–7(–12)-leaved stems. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Although Draba carnosula was reduced by Hitchcock to a variety of D. howellii, it differs significantly by having leafless flowering stems, 2–6-flowered racemes, and broadly winged seeds 3–4.5 mm in diam. Draba howellii almost always has 1–3-leaved stems, (5–)7–18(–25)-flowered racemes, and not winged, distally appendaged seeds 1–1.6 mm in diam. Draba carnosula is known only from a few collections from Mount Eddy in Trinity County and the north side of Mount Shasta in Siskiyou County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 314. | FNA vol. 7, p. 299. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. hirta var. tenella, D. kananaskis, D. longipes | D. howellii var. carnosula |
Name authority | Komarov: Repert. Spec. Nov. Regni Veg. 13: 167. (1914) | O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 82. (1927) |
Web links |