Draba incerta |
Draba santaquinensis |
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Biennials or perennials; (short-lived); caudex simple (poorly developed, without persistent leaf bases); not scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
usually branched, 1.1–3.4 dm, hirsute proximally, trichomes mostly simple and 2-rayed, 0.2–1.5 mm, pubescent distally, trichomes mostly 2- or 3-rayed, 0.2–1 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiole (obscure), ciliate, (trichomes simple and 2-rayed); blade obovate to oblanceolate, 1.6–2.5 cm × 5–10 mm, margins usually entire, rarely denticulate, surfaces pubescent, abaxially with stalked, mostly cruciform trichomes, 0.5–1 mm, adaxially with simple and stalked, 2–4-rayed ones. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
usually 1–5; sessile or subsessile; blade oblanceolate to oblong, margins often denticulate, (ciliate proximally, with 2- or 3-rayed trichomes). |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
9–25-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent, trichomes 2- or 3-rayed. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals oblong-obovate, 2–3.5 mm, pubescent, (trichomes short- to long-stalked, 2–4-rayed, 0.2–0.8 mm); petals yellow (often fading whitish), oblanceolate, 3–5 × 1–1.5 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
horizontal to divaricate-ascending, usually straight, rarely curved upward, 7–10 mm, pubescent, trichomes 2- or 3-rayed. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
narrowly elliptic to slightly falcate, plane, flattened, (7–)10–16 × (2–)2.5–3.7 mm; valves pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.5 mm; ovules 14–22 per ovary; style 0.9–1.2 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
oblong, 1.1–1.5 × 0.7–0.9 mm. |
2n | = 112. |
= 20. |
Draba incerta |
Draba santaquinensis |
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Phenology | Flowering Jun–Aug. | Flowering late Apr-early Jun. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Limestone outcrops and rocky slopes in mixed conifer communities |
Elevation | 0-3300 m (0-10800 ft) | 1800-2400 m (5900-7900 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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UT |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although Draba santaquinensis was included within D. brachystylis by previous authors, I. A. Al-Shehbaz and M. D. Windham (2007) have shown that it is distinct both morphologically and chromosomally. It is currently known only from Utah County (American Fork, Provo, and Santaquin canyons) in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 334. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | Windham & Allphin: Harvard Pap. Bot. 12: 410. (2007) |
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