Draba incerta |
Draba reptans |
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
Carolina Draba, Carolina whitlow-grass, Carolina whitlow-mustard, creeping Draba |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Annuals; scapose or subscapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
(simple or few from, or distal to, base), unbranched, (0.1–)0.3–1.2(–1.6) dm, sparsely to densely pubescent proximally, trichomes 2 (or 3)-rayed, 0.1–0.6 mm, sometimes with simple or spurred ones, to 0.9 mm, distally usually glabrous, rarely with few trichomes. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
not rosulate; petiole obscure; blade elliptic or spatulate to obovate or suborbicular, 0.5–2.3(–3) cm × 1.5–8(–13) mm, margins entire, surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, adaxially with simple trichomes, 0.6–1 mm, with stalked, 2-rayed ones, to 0.7 mm. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
usually 0, rarely 1–3; sessile; blade similar to basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
(3–)5–12(–16)-flowered, ebracteate, (subumbellate), not elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
(late-season ones cleistogamous, apetalous); sepals oblong, 1.5–2.3 mm, pubescent, (trichomes simple); petals (rarely absent), white, spatulate, 2–4.5 × 1–1.5 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
horizontal to divaricate-ascending, straight, (1–)2–7(–9) mm, glabrous or glabrate. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
linear to linear-oblong, plane, flattened, (5–)7–16(–20) × 1.2–2.3 mm; valves glabrous or pubescent, trichomes usually simple and antrorse, 0.1–0.3 mm, rarely with fewer, spurred or 2-rayed ones; ovules 32–88 per ovary; style 0.02–0.1 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
oblong to ovoid, 0.5–0.8 × 0.3–0.5 mm. |
2n | = 112. |
= 16, 30, 32. |
Draba incerta |
Draba reptans |
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Phenology | Flowering Jun–Aug. | Flowering Feb–Jun(-Aug). |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rock outcrops, dry slopes and hillsides, prairies, glades, roadsides, disturbed sites |
Elevation | 0-3300 m (0-10800 ft) | 0-3000 m (0-9800 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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AL; AR; AZ; CA; CO; GA; IA; ID; IL; KS; MA; MI; MN; MO; MT; NC; ND; NE; NM; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
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Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba reptans is often confused with D. cuneifolia, but the two are easily separated. The rachises and pedicels of D. reptans are usually glabrous (rarely with a few isolated trichomes); those of D. cuneifolia are always densely pubescent. Interestingly, both species show parallel variations in chromosome number; it is currently unclear whether this variation is real or the result of misidentified specimens and/or erroneous counts. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 333. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | Arabis reptans, D. caroliniana, D. caroliniana var. dolichocarpa, D. caroliniana var. hunteri, D. caroliniana var. micrantha, D. caroliniana subsp. stellifera, D. caroliniana var. umbellata, D. coloradensis, D. hispidula, D. micrantha, D. reptans var. micrantha, D. reptans subsp. stellifera, D. reptans var. stellifera, D. umbellata, Tomostima caroliniana, Tomostima hispidula, Tomostima micrantha |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | (Lamarck) Fernald: Rhodora 36: 368. (1934) |
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