Draba incerta |
Draba pedicellata |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
Cusick's Draba, rocky-slope Draba, rocky-slope whitlow-grass, stalk whitlow-cress |
|
Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (loosely cespitose); caudex branched (with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, (0.15–)0.3–1.3(–1.6) dm, pubescent proximally, glabrous or nearly so distally, trichomes 2- or 3-rayed and simple, 0.1–1 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; subsessile; blade oblanceolate to obovate, 0.4–1.6(–2) cm × 2–4(–6) mm, margins entire, (not ciliate), surfaces pubescent with stalked, 2–4-rayed trichomes, 0.2–0.9 mm, rarely adaxially with simple trichomes, (midvein obscure abaxially). |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
0. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
3–20(–27)-flowered, ebracteate, slightly to considerably elongated in fruit; rachis straight or flexuous, usually glabrous, rarely pubescent, trichomes simple and 2–4-rayed. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals broadly ovate, 1.8–2.5 mm, glabrous or pubescent, (trichomes simple and branched); petals yellow, spatulate, 3.5–5.5 × 1.5–2.2 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
divaricate to horizontal, straight or curved upward (expanded basally or not), 4–9(–13) mm, glabrous or pubescent, trichomes simple and 2–4-rayed. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
lanceolate to ellipsoid or ovoid, plane (not curved), inflated or slightly flattened, (4.2–)5.3–10(–13) × 2.5–4.5(–5.5) mm; valves glabrous or pubescent, trichomes simple and short-stalked, 2-rayed, 0.05–0.3 mm; ovules 8–16 per ovary; style 0.3–1(–1.2) mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
oblong, 1.2–1.6 × 0.7–0.9 mm. |
2n | = 112. |
= 20. |
Draba incerta |
Draba pedicellata |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rock outcrops, talus, steep gravel slopes in pinyon-juniper, mixed conifer, and subalpine meadow communities |
Elevation | 0-3300 m (0-10800 ft) | 2300-3800 m (7500-12500 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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NV; UT
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Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba pedicellata is known from Elko, Eureka, Nye, and White Pine counties in northeastern Nevada and to Tooele County in northwestern Utah. It was considered a variety of D. cusickii by R. C. Rollins (1993); chromosome number, morphology, and geographic distribution suggest a closer relationship to D. sphaeroides. For a detailed discussion and comparison of these species, consult M. D. Windham (2004). N. H. Holmgren (2005b) divided D. pedicellata into two varieties: var. pedicellata, which is found on limestone throughout the species range, and var. wheelerensis, which is found on non-calcareous substrates in White Pine County. Although var. wheelerensis has slightly more spreading pedicels on somewhat more flexuous rachises, it appears to be little more than a dwarf alpine form of the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 328. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. cusickii var. pedicellata, D. pedicellata var. wheelerensis |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | (Rollins & R. A. Price) Windham: Madroño 50: 221. (2004) |
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