Draba incerta |
Draba oxycarpa |
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (covered with persistent petiole remains); scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, 0.2–1.1 dm, pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.1–0.5 mm, (some rays spurred). |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiolate; petiole base and margin ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to lanceolate, 0.8–2.2 cm × 2–6 mm, margins often entire, surfaces pubescent abaxially with stalked, 2–5-rayed, stellate trichomes, 0.1–0.4 mm, often some rays spurred and trichomes appearing subdendritic, rarely glabrate, or with simple and 2-rayed ones, (midvein obscure abaxially, not thickened), adaxially glabrous or sparsely pubescent with simple trichomes, sometimes with long-stalked, 2- or 3-rayed ones. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
0. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
(2–)4–13-flowered, ebracteate, (subcorymbose), not elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals (grayish white), ovate to broadly so, 2.5–3 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals creamy white to yellow, broadly obovate, 3.5–5 × 2.5–3.5 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
subhorizontal to divaricate-ascending (not appressed to rachis), often slightly curved upward, 4–7(–10) mm, pubescent, trichomes simple (0.4–1 mm), and 2–4-rayed (0.1–0.4 mm). |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
ovate to ovate-oblong, plane, flattened, 5–10 × 3–5 mm; valves puberulent along replum, trichomes simple, 0.1–0.3 mm; ovules (16–)18–28 per ovary; style 0.2–0.6 mm (stigma as wide as style). |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
(black), ovoid, 1–1.3 × 0.6–0.8 mm. |
2n | = 112. |
= 64. |
Draba incerta |
Draba oxycarpa |
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Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rocky soils |
Elevation | 0-3300 m (0-10800 ft) | |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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Greenland; Europe (Norway [including Spitsbergen], Sweden); Atlantic Islands (Iceland) |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba oxycarpa is an octoploid that most closely resembles the decaploid D. alpina. It is distinguished from that species by having creamy white to pale yellow (versus bright yellow) petals, gray-green (versus purplish tinged) sepals, petals 2.5–3.5 (versus 1.7–2.5) mm wide, stems pubescent proximally with primarily branched (versus primarily simple) trichomes, nearly black (versus pale brown) seeds, and ovate to ovate-oblong (versus elliptic) fruits 3–5 (versus 2–3) mm wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 325. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. alpina var. oxycarpa, D. gredinii |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | Sommerfelt: Mag. Naturvidensk., n. s. 1: 240. (1833) |
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