Draba incerta |
Draba nivalis |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
snow Draba, snow whitlow-grass, yellow arctic Draba |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose, usually forming mats); caudex branched (covered with persistent leaf bases); usually scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, 0.2–0.8(–1.2) dm, pubescent throughout, trichomes minutely stalked, 8–15-rayed, stellate, (non-crisped), 0.03–0.15 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiole (obsolete), ciliate throughout; blade oblanceolate to obovate, 0.2–0.9(–1.5) cm × 1–3(–5) mm, margins entire, (not ciliate), surfaces usually pubescent, rarely glabrescent, with short-stalked, 8–15-rayed, stellate, (non-crisped) trichomes, 0.05–0.15 mm. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
0 or 1; sessile; blade ovate or oblong, margins entire, surfaces pubescent as basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
3–9(–11)-flowered, ebracteate, elongated in fruit; rachis often slightly flexuous, pubescent as stem. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and short-stalked, 2–5-rayed); petals white, spatulate to oblanceolate, 2–3.5 × 0.8–1.4 mm; anthers ovate, 0.2–0.3 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
usually ascending, rarely divaricate, straight, 1–4.5(–8) mm, pubescent as stem. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
elliptic to narrowly oblong-elliptic, twisted or not, flattened, 3.5–9 × 1.5–2.2 mm; valves usually glabrous, rarely with 3–6-rayed trichomes on replum; ovules 12–24(–28) per ovary; style 0.1–0.4 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
ovoid, 0.6–1 × 0.5–0.6 mm. |
2n | = 112. |
= 16. |
Draba incerta |
Draba nivalis |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rock outcrops and fellfields, meadows, open tundra, stream banks, gravelly beaches, roadsides |
Elevation | 0-3300 m (0-10800 ft) | 0-2500 m (0-8200 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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AK; AB; BC; MB; NF; NT; NU; ON; QC; YT; Greenland; Europe (Finland, Norway [including Svalbard], n, w Sweden); e Asia (Russian Far East, n Siberia); Atlantic Islands (Iceland); circumpolar |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba nivalis is most often confused with D. lonchocarpa, and C. L. Hitchcock (1941) reduced the latter to a variety (var. elongata) of D. nivalis. A critical comparison reveals that there are enough differences to warrant their recognition as distinct species. Draba nivalis is distinguished by having non-ciliate leaf bases, basal leaf blades pubescent with trichomes 0.05–0.15 mm, stems and pedicels that are always pubescent, and fruits that are elliptic to narrowly elliptic-oblong, plane, and 3.5–9 mm. By contrast, D. lonchocarpa has ciliate leaf bases, basal leaf blades pubescent with trichomes 0.15–0.5 mm, stems and pedicels that are glabrous or sparsely pubescent, and fruits that are linear to lanceolate or oblong, slightly twisted or plane, and 6–15(–18) mm. Except for Alaska, D. nivalis does not grow in the continental United States, whereas D. lonchocarpa grows in nearly all Mountain and Pacific states. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 322. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. caesia |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | Liljeblad: Utkast Sv. Fl., 236. (1792) |
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