Draba incerta |
Draba macounii |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
Macoun's Draba, Macoun's whitlow-grass |
|
Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex simple or branched (with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, (0.06–)0.1–0.4(–0.6) dm, usually pubescent, rarely glabrous, trichomes simple and stalked, 2–4-rayed, 0.15–0.9 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiole (obscure), margin ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate, (0.4–)0.6–1(–1.5) cm × (1–)2–4 mm, margins entire, surfaces usually pubescent abaxially, rarely glabrous, with short-stalked, 2–5-rayed trichomes, 0.1–0.5 mm, adaxially glabrous or pubescent with simple and short-stalked, 2-rayed trichomes. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
0. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
3–10(–13)-flowered, ebracteate, (subumbellate), slightly elongated in fruit; rachis not flexuous, usually pubescent as stem, rarely glabrous. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple and short-stalked, 2- or 3-rayed); petals pale yellow, spatulate, 2.7–4 × 1–2 mm; anthers ovate, 0.2–0.3 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
horizontal to divaricate (not expanded basally), curved upward, 1.5–4.5(–6) mm, abaxially pubescent as stem, adaxially usually glabrous, rarely throughout. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
subglobose to ovoid or ellipsoid, plane, inflated at least basally, 4–8 × 2–4.5 mm; valves glabrous; ovules 10–14(–16) per ovary; style 0.1–0.5 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
ovoid, 1–1.2 × 0.6–0.8 mm. |
2n | = 112. |
= 64. |
Draba incerta |
Draba macounii |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rock outcrops, talus, tundra |
Elevation | 0-3300 m (0-10800 ft) | 700-2800 m (2300-9200 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
|
AK; MT; AB; BC; YT |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba macounii is often confused with the circumpolar decaploid (2n = 80) D. alpina. Plants of the former have scapes (0.6–)1–4(–6) cm, fruiting pedicels divaricate to horizontal, gently curved, and 1.5–4.5(–6) mm, and petals pale yellow, spatulate, and 2.7–4 × 1–2 mm. By contrast, D. alpina has scapes (3–)5–17(–28) cm, fruiting pedicels divaricate-ascending to ascending, often straight, and (3–)4–14(–30) mm, and petals bright yellow, obovate, and 3.5–5 × 1.7–2.5 mm. R. C. Rollins (1993) indicated that the species occurs in Colorado, but we have not seen any material from that state. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 318. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 97. (1927) |
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