Flora of North America species comparison

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Draba incerta

Draba lactea

whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort

milky Draba, milky whitlow-grass
Habit Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. Perennials; (cespitose); caudex branched (with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.
Stems

unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones).

unbranched, 0.2–1.1(–1.5) dm,glabrous throughout or sparsely pubescent proximally, trichomes short-stalked, substellate, 2–8-rayed, (non-crisped), 0.5–0.3 mm.
Basal leaves

rosulate; petiolate;

petiole (0–1 cm), ciliate throughout;

blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially.

rosulate; petiolate;

petiole (persistent, strongly thickened), margin usually ciliate, (trichomes usually simple and 2-rayed, 0.3–1 mm);

blade oblanceolate to obovate, (0.3–)0.5–1.1(–1.7) cm × (1–)2–6 mm, margins usually entire, rarely denticulate, (sometimes ciliate), surfaces sometimes pubescent with stellate to subdendritic, 4–12-rayed, (non-crisped) trichomes, 0.1–0.4 mm, (midvein persistent, prominent, strongly thickened).
Cauline leaves

usually 0 (or 1, as a bract);

sessile;

blade linear to oblong, margins entire, surfaces pubescent as basal.

0 (or, rarely, 1 as a bract).
Racemes

3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit;

rachis not flexuous, glabrous or pubescent as stem.

2–8(–12)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, usually glabrous, rarely sparsely pubescent as stem basally.
Flowers

sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed);

petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm;

anthers ovate, 0.3–0.5 mm.

sepals ovate, 1.8–3 mm, usually glabrous, rarely sparsely pubescent subapically, (trichomes simple);

petals white, obovate, 3–5 × 1.8–3 mm;

anthers ovate, 0.3–0.4 mm.
Fruiting pedicels

ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate.

divaricate-ascending, straight, (1–)2–5(–10) mm, glabrous.
Fruits

broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm;

valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm;

ovules 8–16(–20) per ovary;

style 0.2–0.9 mm.

oblong to elliptic-lanceolate or ovate to broadly so, plane, flattened, 4–8 × (1.5–)2–3 mm;

valves glabrous;

ovules (10–)14–22(–26) per ovary;

style 0.1–0.4 mm.
Seeds

oblong, 1.1–1.5 × 0.7–1 mm.

ovoid, 0.8–1.1 × 0.5–0.6 mm.
2n

= 112.

= 32, 48.

Draba incerta

Draba lactea
Phenology Flowering Jun–Aug. Flowering Jun–Aug.
Habitat Rock outcrops, talus, gravelly areas, tundra Rock outcrops, talus, rocky hillsides and ridges, open gravelly areas, seepage swales, meadows
Elevation 0-3300 m (0-10800 ft) 0-2000 m (0-6600 ft)
Distribution
from FNA
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NL; NT; NU; QC; YT; Greenland; Europe (n Finland, Norway, w Sweden); Asia (Russian Far East, c, n Siberia); Atlantic Islands (Iceland); circumpolar
[BONAP county map]
Discussion

Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta.

Draba incerta is found near sea level in Alaska.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

T. W. Böcher (1966) postulated that Draba lactea originated from hybridization between D. fladnizensis and D. nivalis, but A.-C. Scheen et al. (2002) showed that it is more closely allied to D. subcapitata. By contrast, H. H. Grundt et al. (2004) concluded that hexaploid D. lactea originated from tetraploids of the same species, which in turn originated from the diploid D. palanderiana lineage. They suggested that D. lactea probably originated multiple times in the Beringian area and migrated to reach its present circumpolar distribution. The hexaploids are distributed throughout the species range, whereas the tetraploids are known only from Alaska and the Russian Far East (Grundt et al. 2005b).

Draba fernaldiana, which was collected from Southampton Island (Nunavut), was not mentioned by R. C. Rollins (1993). The plants are completely glabrous except for leaf margins, which are ciliate with simple and sparse 2-rayed trichomes. The taxon resembles some forms of D. lactea and is tentatively herein included within that species. The only conflict in such placement is petal color, which was listed in the original description of D. fernaldiana as pale yellow instead of white.

Glabrous or glabrescent forms of Draba lactea are quite common in the Canadian Arctic Archipelago, whereas pubescent forms predominate in Alaska and the Russian Far East.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 7, p. 313. FNA vol. 7, p. 316.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei D. allenii, D. boecheri, D. fernaldiana
Name authority Payson: Amer. J. Bot. 4: 261. (1917) Adams: Mém. Soc. Imp. Naturalistes Moscou 5: 104. (1817)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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