Draba incerta |
Draba inexpectata |
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex simple or branched (with persistent leaf remains); not scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, 0.1–0.4 dm, pubescent, trichomes 5–10-rayed, 0.05–0.2 mm, and fewer, simple and 2-rayed ones, 0.3–0.6 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; subsessile; petiole ciliate; blade obovate to oblanceolate, 0.2–0.4 cm × 0.7–1.5 mm, (base sparsely ciliate, trichomes simple and 2-rayed, 0.4–0.6 mm), margins entire, surfaces densely pubescent with subsessile, 6–12-rayed, stellate trichomes, 0.25–0.7 mm. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
1–3 (or 4); sessile; blade ovate to lanceolate, margins entire or denticulate, surfaces pubescent as basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
2–5(–8)-flowered, proximalmost 1–3 flowers bracteate, slightly elongated in fruit; rachis not flexuous, pubescent, trichomes 5–10-rayed. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals (sometimes persistent), ovate, 1.2–1.7 mm, pubescent, (trichomes simple and branched); petals white, obovate, 2–2.5 × 1–1.2 mm; anthers ovate, 0.1–0.2 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
ascending, straight, 0.7–2(–3) mm, pubescent as rachis. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
elliptic to lanceolate, slightly twisted or plane, flattened, 3.5–5 × 1.5–2 mm; valves pubescent, trichomes short-stalked, (3 or) 4(–6)-rayed, 0.1–0.2 mm; ovules 16–24 per ovary; style 0.2–0.4 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
oblong, 0.7–1 × 0.4–0.5 mm. |
2n | = 112. |
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Draba incerta |
Draba inexpectata |
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Phenology | Flowering Jun–Aug. | Flowering Jul–Aug. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rocky ridges and slopes, on accumulated soil among boulders, subalpine fir and juniper communities |
Elevation | 0-3300 m (0-10800 ft) | 3100-3700 m (10200-12100 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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UT |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba inexpectata is known from the Uinta Mountains in Summit County. The measurements above are based on the holotype (Franklin 6328) and two of the three paratypes (Franklin 6293, 6331), all at BRY. One paratype (Goodrich 26166, BRY) is a robust, completely sterile plant of uncertain identity and was not used for the description above. Although compared in the original description with Draba lonchocarpa (with which it grows sympatrically), D. inexpectata appears most closely related to D. cana. Draba inexpectata is distinguished from the latter by having stems 1–4 cm, basal leaves non-ciliate, racemes that are 2–5(–8)-flowered and scarcely elongated in fruit, fruits elliptic to lanceolate, 3.5–5 mm, and ovules 16–24 per ovary. By contrast, D. cana has stems (6–)10–30 cm, basal leaves ciliate at the apex, racemes that are (10–)15–47(–63)-flowered and often considerably elongated in fruit, fruits linear-lanceolate or very rarely ovate-oblong, (5–)6–11 mm, and ovules 28–48 per ovary. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 313. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | S. L. Welsh: in S. L. Welsh et al., Utah Fl. ed. 3, 272. (2003) |
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