Draba incerta |
Draba howellii |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
Howell's Draba, Howell's whitlow-grass, rosette Draba |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (loosely cespitose); caudex branched (somewhat surculose, with persistent leaf remains, branches sometimes terminating in sterile rosettes); often scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, (0.2–)0.4–1.1(–1.5) dm, usually pubescent throughout, rarely glabrous, trichomes simple and 2–4-rayed, 0.1–0.6 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; blade (somewhat fleshy), oblanceolate or spatulate to obovate, 0.4–1.6(–2.5) cm × (1.5–)3–6(–10) mm, margins entire, surfaces pubescent with stalked, cruciform, and fewer 2- or 3-rayed trichomes, 0.07–0.5 mm, rarely both surfaces glabrous and trichomes on margins, (midvein obscure abaxially). |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
0–3 (or 4); sessile; blade ovate to oblong, margins entire, pubescent as basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
(5–)7–18(–25)-flowered, usually ebracteate, sometimes proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, usually pubescent as stem, rarely glabrous. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 2.5–3.2 mm, glabrous or pubescent, (trichomes 2–4-rayed); petals yellow, oblanceolate, 5.5–8 × 1–2 mm; anthers oblong, 0.7–0.9 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
divaricate-ascending, straight, (4–)7–10 mm, usually pubescent as stem, rarely glabrous. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
lanceolate to elliptic-lanceolate or broadly ovate, plane (not curved), strongly flattened, 6–11(–15) × 3–5 mm; valves usually pubescent, rarely glabrous, trichomes simple and 2–4-rayed, 0.05–0.3 mm; ovules 8–22 per ovary; style (0.7–)1.6–3 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
oblong, 1–1.6 × 0.8–1 mm, (sometimes distally appendaged). |
2n | = 112. |
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Draba incerta |
Draba howellii |
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Phenology | Flowering Jun–Aug. | Flowering Jul–Aug. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rocky summits, cracks in granite walls, rock crevices |
Elevation | 0-3300 m (0-10800 ft) | 1900-2700 m (6200-8900 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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CA; OR |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba howellii is known from Siskiyou and Trinity counties, California, and Josephine County, Oregon. One collection, Tracy 14623 (DS, GH, UC), is unusual in having glabrous stems, pedicels, and leaf blade surfaces. In this regard, it resembles D. carnosula, but in all other respects (bracts, seeds, inflorescences, etc.), it is indistinguishable from D. howellii. For characteristics distinguishing the two species, see 23. D. carnosula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 312. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | S. Watson: Proc. Amer. Acad. Arts 20: 354. (1885) |
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