FNA: Draba incerta vs. Draba hitchcockii

	Draba incerta	Draba hitchcockii
	whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort	lost river Draba
Habit	Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose.	Perennials; (densely cespitose); caudex branched (densely covered with persistent leaves, branches sometimes terminating in sterile rosettes); scapose.
Stems	unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones).	unbranched, (0.1–)0.3–1(–1.3) dm, hirsute throughout, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.6 mm.
Basal leaves	rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially.	(densely imbricate); rosulate; sessile; blade narrowly oblanceolate to oblong-linear, 0.3–1.2(–1.5) cm × 1–2 mm, margins entire, (ciliate, trichomes simple, 0.3–1.2 mm), surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.08–0.45 mm, (midvein obscure), adaxially glabrous proximally, sparsely pubescent distally with mostly simple trichomes.
Cauline leaves	usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal.	0.
Racemes	3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem.	4–15-flowered, ebracteate, elongated in fruit; rachis not flexuous, hirsute as stem.
Flowers	sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm.	sepals oblong, 2.5–3.5 mm, hirsute, (trichomes simple and stalked, 2–4-rayed); petals white, obovate, 5–6.5 × 2–3.5 mm; anthers ovate, 0.5–0.6 mm.
Fruiting pedicels	ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate.	ascending, straight, (2–)4–13(–18) mm, hirsute as stem.
Fruits	broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm.	ovate to broadly oblong or elliptic, plane, flattened, (3–)4–7(–10) × (2.5–)3.5–5 mm; valves pubescent, trichomes simple and short-stalked, 2(–4)-rayed, 0.06–0.5 mm; ovules 8–12 per ovary; style (0.8–)1–1.7(–2) mm.
Seeds	oblong, 1.1–1.5 × 0.7–1 mm.	oblong, 1.4–1.8 × 0.8–1 mm.
2n	= 112.	= 54.

	Draba incerta	Draba hitchcockii
Phenology	Flowering Jun–Aug.	Flowering May–Jun.
Habitat	Rock outcrops, talus, gravelly areas, tundra	Limestone outcrops and gravelly soil
Elevation	0-3300 m (0-10800 ft)	1800-2200 m (5900-7200 ft)
Distribution	AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT [WildflowerSearch map] [BONAP county map]	ID [BONAP county map]
Discussion	Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Draba hitchcockii is known from the Lost River Range in Butte and Custer counties. Based on morphological and chromosomal evidence, M. D. Windham (2004) suggested that it may be an allopolyploid resulting from hybridization between D. oreibata and D. paysonii. Draba hitchcockii is superficially similar to D. jaegeri, a taxon known from the Charleston Mountains of Clark County, Nevada. Both are cespitose perennials with relatively large, white flowers and a chromosome number (2n = 54) otherwise unknown in Draba (Windham). Features distinguishing these two taxa are provided in the discussion of 51. D. jaegeri. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 7, p. 313.	FNA vol. 7, p. 311.
Parent taxa	Brassicaceae > tribe Arabideae > Draba	Brassicaceae > tribe Arabideae > Draba
Sibling taxa	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms	D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei
Name authority	Payson: Amer. J. Bot. 4: 261. (1917)	Rollins: J. Arnold Arbor. 64: 500. (1983)
Web links	Local floras: BC, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local Web sites: PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Draba hitchcockii

whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort

lost river Draba

Habit

Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose.

Perennials; (densely cespitose); caudex branched (densely covered with persistent leaves, branches sometimes terminating in sterile rosettes); scapose.

Stems

unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones).

unbranched, (0.1–)0.3–1(–1.3) dm, hirsute throughout, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.6 mm.

Basal leaves

rosulate; petiolate;

petiole (0–1 cm), ciliate throughout;

blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially.

(densely imbricate); rosulate;

sessile;

blade narrowly oblanceolate to oblong-linear, 0.3–1.2(–1.5) cm × 1–2 mm, margins entire, (ciliate, trichomes simple, 0.3–1.2 mm), surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.08–0.45 mm, (midvein obscure), adaxially glabrous proximally, sparsely pubescent distally with mostly simple trichomes.

Cauline leaves

usually 0 (or 1, as a bract);

sessile;

blade linear to oblong, margins entire, surfaces pubescent as basal.

Racemes

3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit;

rachis not flexuous, glabrous or pubescent as stem.

4–15-flowered, ebracteate, elongated in fruit;

rachis not flexuous, hirsute as stem.

Flowers

sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed);

petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm;

anthers ovate, 0.3–0.5 mm.

sepals oblong, 2.5–3.5 mm, hirsute, (trichomes simple and stalked, 2–4-rayed);

petals white, obovate, 5–6.5 × 2–3.5 mm;

anthers ovate, 0.5–0.6 mm.

Fruiting pedicels

ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate.

ascending, straight, (2–)4–13(–18) mm, hirsute as stem.

Fruits

broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm;

valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm;

ovules 8–16(–20) per ovary;

style 0.2–0.9 mm.

ovate to broadly oblong or elliptic, plane, flattened, (3–)4–7(–10) × (2.5–)3.5–5 mm;

valves pubescent, trichomes simple and short-stalked, 2(–4)-rayed, 0.06–0.5 mm;

ovules 8–12 per ovary;

style (0.8–)1–1.7(–2) mm.

Seeds

oblong, 1.1–1.5 × 0.7–1 mm.

oblong, 1.4–1.8 × 0.8–1 mm.

= 112.

= 54.

Draba incerta

Draba hitchcockii

Phenology

Flowering Jun–Aug.

Flowering May–Jun.

Habitat

Rock outcrops, talus, gravelly areas, tundra

Limestone outcrops and gravelly soil

Elevation

0-3300 m (0-10800 ft)

1800-2200 m (5900-7200 ft)

Distribution

AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT

[WildflowerSearch map]

[BONAP county map]

Discussion

Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta.

Draba incerta is found near sea level in Alaska.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba hitchcockii is known from the Lost River Range in Butte and Custer counties. Based on morphological and chromosomal evidence, M. D. Windham (2004) suggested that it may be an allopolyploid resulting from hybridization between D. oreibata and D. paysonii. Draba hitchcockii is superficially similar to D. jaegeri, a taxon known from the Charleston Mountains of Clark County, Nevada. Both are cespitose perennials with relatively large, white flowers and a chromosome number (2n = 54) otherwise unknown in Draba (Windham). Features distinguishing these two taxa are provided in the discussion of 51. D. jaegeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 7, p. 313.

FNA vol. 7, p. 311.

Parent taxa

Brassicaceae > tribe Arabideae > Draba

Sibling taxa

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

Synonyms

D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei

Name authority

Payson: Amer. J. Bot. 4: 261. (1917)

Rollins: J. Arnold Arbor. 64: 500. (1983)

Web links

Local floras: BC, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Draba incerta

Draba hitchcockii

Draba incerta

Draba hitchcockii