Draba incerta |
Draba grayana |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
Gray's Draba |
|
Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (densely cespitose); caudex branched (covered with persistent, somewhat thickened, dry petioles); not scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, 0.08–0.6 dm, densely to moderately pubescent throughout, trichomes simple and stalked, 2- or 3-rayed, 0.2–1 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiolate; petiole ciliate, (trichomes simple, 0.2–1 mm); blade linear to linear-oblanceolate, (0.4–)0.6–1.5(–2) cm × 1–2 mm, margins entire, (pubescent as petiole), surfaces pubescent abaxially with simple and stalked, 2- or 3-rayed trichomes, 0.1–0.4 mm, adaxially glabrate or subapically sparsely pubescent with simple trichomes. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
1–4; sessile; blade linear to linear-oblanceolate, margins entire, surfaces pubescent as basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
5–12(–16)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and short-stalked, 2- or 3-rayed); petals bright yellow, spatulate, 3–4.5 × 1–2 mm, (flared, clawed); anthers ovate, 0.3–0.5 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
divaricate-ascending, straight or slightly curved upward, 1.5–6 mm, pubescent as stem. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
lanceolate to oblong-lanceolate, slightly twisted or plane, flattened, 4–12 × 1.5–3 mm; valves glabrous or pubescent, trichomes simple, 0.07–0.2 mm; ovules 16–20 per ovary; style 0.4–1.2 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
ovoid, 1–1.4 × 0.7–1 mm. |
2n | = 112. |
= 24 ± 3. |
Draba incerta |
Draba grayana |
|
Phenology | Flowering Jun–Aug. | Flowering Jul–Aug. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Alpine tundra, fellfields, gravelly wet meadows |
Elevation | 0-3300 m (0-10800 ft) | 3500-4000 m (11500-13100 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
|
CO |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba grayana is an apomictic species clearly related to D. exunguiculata, with which it is occasionally sympatric (see discussion of 36. D. exunguiculata for distinguishing features). It is known from Chaffe, Clear Creek, Larimer, Park, and Summit counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 310. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. streptocarpa var. grayana, D. alpicola |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | (Rydberg) C. L. Hitchcock: Revis. Drabas W. N. Amer., 29. (1941) |
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