Draba incerta |
Draba exunguiculata |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
clawless Draba |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (densely cespitose); caudex branched (covered with persistent, somewhat thickened, dry petioles); not scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, 0.15–0.7 dm, glabrous throughout or sparsely pubescent, trichomes simple, 0.3–0.9 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiolate; petiole ciliate, (trichomes simple, 0.3–1.1 mm); blade linear to linear-oblanceolate, (0.5–)0.8–2(–2.5) cm × 1–3 mm, margins entire, (pubescent as petiole), surfaces abaxially sparsely pubescent with simple and stalked, 2- or 3-rayed trichomes, 0.1–0.4 mm, adaxially glabrous or subapically sparsely pubescent with simple trichomes. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
1–4; sessile; blade linear to linear-oblanceolate, margins entire, surfaces pubescent as basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
4–13(–20)-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent, trichomes simple, to 1 mm. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 2–2.5 mm, pubescent, (trichomes simple); petals (erect), yellow, obovate, 2–2.5(–3) × 1.5–2 mm, (not clawed); anthers oblong, 0.6–0.9 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
divaricate-ascending, straight or slightly curved upward, 1–6 mm, usually glabrous, rarely with few, simple trichomes. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
lanceolate to oblong-lanceolate, plane, flattened, 5–13 × 1.5–3 mm; valves glabrous; ovules 20–24 per ovary; style 0.3–1 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
ovoid, 1–1.3 × 0.6–0.8 mm. |
2n | = 112. |
= 56 ± 5. |
Draba incerta |
Draba exunguiculata |
|
Phenology | Flowering Jun–Aug. | Flowering Jul–Aug. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Open knolls, talus, gravelly alpine slopes, rocky alpine tundra |
Elevation | 0-3300 m (0-10800 ft) | 3600-4300 m (11800-14100 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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CO |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba exunguiculata occupies alpine areas of Clear Creek, El Paso, Grand, and Summit counties in central Colorado. Draba grayana is found in the same general area; both species are apomicts that occasionally grow sympatrically. Draba exunguiculata is easily distinguished from D. grayana by having stems and fruiting pedicels glabrous or sparsely pubescent with simple trichomes (versus moderately to densely pubescent with simple and 2- or 3-rayed trichomes), and by having erect, non-clawed petals, 2–3 mm and slightly longer than sepals (versus flared, clawed petals 3–4.5 mm and nearly twice as long as sepals). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 306. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. chrysantha var. exunguiculata, D. chrysantha |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | (O. E. Schulz) C. L. Hitchcock: Revis. Drabas W. N. Amer., 46. (1941) |
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