FNA: Draba incerta vs. Draba cyclomorpha

	Draba incerta	Draba cyclomorpha
	whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort	granite Draba, Lemmon's Draba
Habit	Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose.	Perennials; (cespitose, not pulvinate); caudex branched (with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.
Stems	unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones).	unbranched, 0.1–0.6(–1) dm, (thinly) pilose throughout, trichomes simple and stalked, 2- (or 3-)rayed, (often crisped), 0.1–0.8 mm.
Basal leaves	rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially.	rosulate; shortly petiolate; petiole base and margin ciliate, (trichomes simple, 0.2–1 mm); blade (somewhat fleshy), oblanceolate to obovate, 0.4–1 cm × 2–5 mm, margins entire, surfaces glabrous or hirsute, abaxially usually with stalked, 2-rayed trichomes, 0.1–0.6 mm, rarely with fewer, simple ones, (midvein obscure), adaxially with almost exclusively simple trichomes, to 0.6 mm.
Cauline leaves	usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal.	0.
Racemes	3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem.	5–14(–20)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem.
Flowers	sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm.	sepals ovate, 1.5–2.2 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, spatulate, 4–5.5 × 1.5–2.5 mm; anthers oblong, 0.6–0.7 mm.
Fruiting pedicels	ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate.	horizontal to divaricate-ascending, often curved upward or straight, 3–8(–10) mm, pilose as stem.
Fruits	broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm.	oblong to oblong-ovate, plane, flattened, 3.5–9 × 2.5–4 mm; valves glabrous; ovules 8–12 per ovary; style 0.3–1 mm.
Seeds	oblong, 1.1–1.5 × 0.7–1 mm.	ovoid, 1.4–1.9 × 1–1.2 mm.
2n	= 112.

	Draba incerta	Draba cyclomorpha
Phenology	Flowering Jun–Aug.	Flowering Jul–Aug.
Habitat	Rock outcrops, talus, gravelly areas, tundra	Basaltic talus, stony ridges
Elevation	0-3300 m (0-10800 ft)	2500-3100 m (8200-10200 ft)
Distribution	AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT [WildflowerSearch map] [BONAP county map]	OR [BONAP county map]
Discussion	Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Draba cyclomorpha was treated by Schulz, C. L. Hitchcock (1941), and R. C. Rollins (1993) as a variety of D. lemmonii. The two taxa are quite distinct morphologically and separated by nearly 1600 kilometers, justifying their treatment as distinct species (I. A. Al-Shehbaz and M. D. Windham 2007). Draba cyclomorpha is known to us only from the Wallowa Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 7, p. 313.	FNA vol. 7, p. 305.
Parent taxa	Brassicaceae > tribe Arabideae > Draba	Brassicaceae > tribe Arabideae > Draba
Sibling taxa	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms	D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei	D. lemmonii var. cyclomorpha
Name authority	Payson: Amer. J. Bot. 4: 261. (1917)	Payson: Amer. J. Bot. 4: 263. (1917)
Web links	Local floras: BC, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Draba cyclomorpha

whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort

granite Draba, Lemmon's Draba

Habit

Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose.

Perennials; (cespitose, not pulvinate); caudex branched (with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.

Stems

unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones).

unbranched, 0.1–0.6(–1) dm, (thinly) pilose throughout, trichomes simple and stalked, 2- (or 3-)rayed, (often crisped), 0.1–0.8 mm.

Basal leaves

rosulate; petiolate;

petiole (0–1 cm), ciliate throughout;

blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially.

rosulate; shortly petiolate;

petiole base and margin ciliate, (trichomes simple, 0.2–1 mm);

blade (somewhat fleshy), oblanceolate to obovate, 0.4–1 cm × 2–5 mm, margins entire, surfaces glabrous or hirsute, abaxially usually with stalked, 2-rayed trichomes, 0.1–0.6 mm, rarely with fewer, simple ones, (midvein obscure), adaxially with almost exclusively simple trichomes, to 0.6 mm.

Cauline leaves

usually 0 (or 1, as a bract);

sessile;

blade linear to oblong, margins entire, surfaces pubescent as basal.

Racemes

3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit;

rachis not flexuous, glabrous or pubescent as stem.

5–14(–20)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed);

petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm;

anthers ovate, 0.3–0.5 mm.

sepals ovate, 1.5–2.2 mm, pubescent, (trichomes simple and short-stalked, 2-rayed);

petals yellow, spatulate, 4–5.5 × 1.5–2.5 mm;

anthers oblong, 0.6–0.7 mm.

Fruiting pedicels

ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate.

horizontal to divaricate-ascending, often curved upward or straight, 3–8(–10) mm, pilose as stem.

Fruits

broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm;

valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm;

ovules 8–16(–20) per ovary;

style 0.2–0.9 mm.

oblong to oblong-ovate, plane, flattened, 3.5–9 × 2.5–4 mm;

valves glabrous;

ovules 8–12 per ovary;

style 0.3–1 mm.

Seeds

oblong, 1.1–1.5 × 0.7–1 mm.

ovoid, 1.4–1.9 × 1–1.2 mm.

= 112.

Draba incerta

Draba cyclomorpha

Phenology

Flowering Jun–Aug.

Flowering Jul–Aug.

Habitat

Rock outcrops, talus, gravelly areas, tundra

Basaltic talus, stony ridges

Elevation

0-3300 m (0-10800 ft)

2500-3100 m (8200-10200 ft)

Distribution

AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT

[WildflowerSearch map]

[BONAP county map]

Discussion

Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta.

Draba incerta is found near sea level in Alaska.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba cyclomorpha was treated by Schulz, C. L. Hitchcock (1941), and R. C. Rollins (1993) as a variety of D. lemmonii. The two taxa are quite distinct morphologically and separated by nearly 1600 kilometers, justifying their treatment as distinct species (I. A. Al-Shehbaz and M. D. Windham 2007). Draba cyclomorpha is known to us only from the Wallowa Mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 7, p. 313.

FNA vol. 7, p. 305.

Parent taxa

Brassicaceae > tribe Arabideae > Draba

Sibling taxa

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

Synonyms

D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei

D. lemmonii var. cyclomorpha

Name authority

Payson: Amer. J. Bot. 4: 261. (1917)

Payson: Amer. J. Bot. 4: 263. (1917)

Web links

Local floras: BC, OR, WA
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Flora of North America species comparison

Draba incerta

Draba cyclomorpha

Draba incerta

Draba cyclomorpha