Draba incerta |
Draba corymbosa |
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
0. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
(brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
2n | = 112. |
= 128, 144. |
Draba incerta |
Draba corymbosa |
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Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces |
Elevation | 0-3300 m (0-10800 ft) | 0-1700 m (0-5600 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 301. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) |
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