Draba incerta |
Draba brachystylis |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
shortstyle Draba |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Annuals or perennials; (short-lived); caudex often simple (poorly developed); not scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched or branched, 0.6–3(–3.7) dm, pubescent throughout, trichomes simple, 0.4–0.9 mm, and 2–4-rayed, 0.05–0.4 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; subsessile; petiole base ciliate, (trichomes simple, 0.3–0.9 mm); blade oblanceolate, 1–3.5 cm × 4–7(–12) mm, margins entire or denticulate, surfaces pubescent, abaxially with stalked, (2–)4-rayed trichomes, 0.1–0.5 mm, adaxially with simple trichomes, 0.2–0.6 mm, with smaller, 2–4-rayed ones. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
(1 or) 2–6(–8); sessile; blade lanceolate or oblanceolate to ovate, margins entire or denticulate, surfaces pubescent as basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
(5–)10–35(–47)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 2–2.7 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals pale yellow (fading white), oblanceolate, 2.5–3.7 × 0.9–1.2 mm; anthers ovate, 0.2–0.3 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
divaricate-ascending, straight or only slightly curved upward, (1.5–)3–8(–10) mm, pubescent throughout, trichomes simple and stalked, 2–4-rayed. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
narrowly elliptic to lanceolate, plane, flattened, (7–)10–16(–19) × 2–3.5 mm; valves pubescent, trichomes simple, spurred, and short-stalked, 2-rayed, 0.05–0.2(–0.3) mm; ovules 20–36(–42) per ovary; style 0.2–0.6(–0.8) mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
oblong, 0.9–1.3 × 0.6–0.7 mm. |
2n | = 112. |
= 44. |
Draba incerta |
Draba brachystylis |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Fir and aspen communities, moist areas on rocky slopes |
Elevation | 0-3300 m (0-10800 ft) | 1700-3000 m (5600-9800 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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UT |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Based on morphological and chromosomal data (L. Allphin and M. D. Windham, unpubl.), Draba brachystylis is hypothesized to be an allopolyploid derived through hybridization between D. albertina and D. santaquinensis. It is known to us only from Cache, Duchesne, Juab, Salt Lake, and Utah counties. C. L. Hitchcock (1941) indicated that the species grows in the Charleston Mountains of Clark County, Nevada, but we have not seen unequivocal material of it from that area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 297. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | Rydberg: Bull. Torrey Bot. Club 29: 240. (1902) |
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