Draba incerta |
Draba argyrea |
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
silver Draba |
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Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; caudex branched (branches loose, creeping, sometimes terminating in sterile shoots); scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, (0.2–)0.3–0.8(–1.1) dm, pubescent proximally, glabrous distally, trichomes short-stalked to subsessile, 5–8-rayed, 0.05–0.2 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; subsessile; petiole base and margin not ciliate, (midvein obscure abaxially); blade oblanceolate to obovate, 0.3–0.7(–1.2) cm × 1–2.5(–4) mm, margins entire, surfaces canescent with short-stalked, 8–12-rayed, stellate trichomes, 0.1–0.2 mm. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
0. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
2–8(–12)-flowered, ebracteate, elongated in fruit; rachis slightly flexuous, usually glabrous, rarely sparsely pubescent as stem. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 2–3 mm, pubescent, (trichomes 4–6-rayed); petals yellow, oblanceolate, 4–6 × 1.5–3 mm; anthers ovate, 0.5–0.6 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
divaricate-ascending, straight or slightly curved upward, (4–)5–12(–19) mm, glabrous. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
lanceolate to elliptic or ovate, plane or slightly twisted, flattened, 5–11 × 2–3.5(–4.5) mm, (symmetric); valves often puberulent, sometimes glabrous, trichomes simple and subsessile, 2–4-rayed, 0.03–0.2 mm; ovules 8–14 per ovary; style (0.7–)1–1.8(–2.4) mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
oblong, 1.2–1.8 × 0.8–1.1 mm. |
2n | = 112. |
= 36. |
Draba incerta |
Draba argyrea |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Jul. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rock crevices, ledges, talus slopes |
Elevation | 0-3300 m (0-10800 ft) | 1000-3100 m (3300-10200 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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ID |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
M. A. Beilstein and M. D. Windham (2003) hypothesized that Draba argyrea is an allopolyploid derived through hybridization between D. lonchocarpa and D. sphaerocarpa. The species is restricted to Blaine, Boise, Custer, and Elmore counties in central Idaho. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 292. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. argyrea var. glabrescens |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | Rydberg: Bull. Torrey Bot. Club 30: 251. (1903) |
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