Draba incerta |
Draba arctogena |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
fellfield whitlowgrass |
|
Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (sometimes with persistent leaf remains); sometimes scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
unbranched, 0.3–1.4 dm, pubescent throughout, trichomes simple, 0.2–1 mm, and 2–6-rayed, 0.05–0.3 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiolate; petiole base and margin ciliate, (trichomes mostly simple, 0.2–1 mm); blade oblanceolate to narrowly obovate, 0.4–1.7 cm × 1.5–6 mm, margins entire or with 1–3 teeth on each side, surfaces pubescent with primarily simple trichomes, 0.3–1mm, abaxially mixed with stalked, 4–10-rayed ones, 0.1–0.2 mm. |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
0 or 1; sessile; blade ovate to oblong, margins entire or toothed, surfaces pubescent as basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
2–14(–17)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 1.8–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals white, spatulate, 3–4 × 1.5–2 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
suberect to ascending, straight, 1–5(–7) mm, pubescent, trichomes simple and 2–4-rayed. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
oblong, plane, flattened, 4–9 × 2–3.5 mm; valves pubescent, trichomes short-stalked, 2–4-rayed, 0.1–0.5 mm, sometimes with simple ones; ovules 20–32 per ovary; style 0.1–0.4 mm (stigma distinctly wider than style). |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
ovoid, 0.7–1 × 0.5–0.6 mm. |
2n | = 112. |
= 48. |
Draba incerta |
Draba arctogena |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Scree, sandy hills, gravel flats |
Elevation | 0-3300 m (0-10800 ft) | 0-450 m (0-1500 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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NT; NU; Greenland |
Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba arctogena is sometimes confused with D. norvegica, from which it differs by having less elongated fruiting racemes, abaxial leaf surfaces with 4–10-rayed (versus cruciform or, rarely, 2–6-rayed) trichomes, and fruits with 2–4-rayed trichomes, 0.1–0.5 mm, (versus glabrous or with simple and 2–4-rayed trichomes, 0.05–0.25 mm). The limits of D. arctica, D. arctogena, D. cinerea, and D. oblongata need to be critically evaluated in light of molecular and cytological data. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 292. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. groenlandica var. arctogena, D. cinerea var. arctogena |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | (E. Ekman) E. Ekman: Svensk Bot. Tidskr. 25: 492. (1932) |
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